AOSA (Association of Official Seed Analysts). 1990. Rules for testing seeds. Journal of Seed Technology, 16, 31–78.

Ahanger M A, Tomar N S, Tittal M, Argal S, Agarwal R M. 2017. Plant growth under water/salt stress: ROS production; antioxidants and significance of added potassium under such conditions. Physiology Molecular Biology of Plants, 23, 731–744.

Ahmad F, Kamal A, Singh A, Ashfaque F, Alamri S, Siddiqui M H, Khan M I R. 2021. Seed priming with gibberellic acid induces high salinity tolerance in Pisum sativum through antioxidants, secondary metabolites and up-regulation of antiporter genes. Plant Biology, 23, 113–121.

Al-Salama Y. 2022. Effect of seed priming with ZnO nanoparticles and saline irrigation water in yield and nutrients uptake by wheat plants. Environmental Sciences Proceedings, 16, 37.

An J, Hu P, Li F, Wu H, Shen Y, White J C, Tian X, Li Z, Giraldo J P. 2020. Emerging investigator series: Molecular mechanisms of plant salinity stress tolerance improvement by seed priming with cerium oxide nano particles. Environmental Science-Nano, 7, 2214–2228.

Arafat Abdel Hamed A L, Abu Alhmad M F, Abdelfattah K E. 2017. The possible roles of priming with ZnO nanoparticles in mitigation of salinity stress in lupine (Lupinus termis) plants. Journal of Plant Growth Regulation, 36, 60–70.

Aragão V P M, Navarro B V, Passamani L Z, Macedo A F, Floh E I S, Silveira V, Santa-Catarina C. 2015. Free amino acids, polyamines, soluble sugars and proteins during seed germination and early seedling growth of Cedrela fissilis Vellozo (Meliaceae), an endangered hardwood species from the Atlantic Forest in Brazil. Theoretical and Experimental Plant Physiology, 27, 157–169.

Ashraf M, McNeilly T. 2004. Salinity tolerance in Brassica oilseeds. Critical Reviews in Plant Sciences, 23, 157–174.

Baldim V, Bedioui F, Mignet N, Margaill I, Berret J F. 2018. The enzyme-like catalytic activity of cerium oxide nanoparticles and its dependency on Ce3+ surface area concentration. Nanoscale, 10, 6971–6980.

Banerjee A, Roychoudhury A. 2021. Role of sugars in mediating abiotic stress tolerance in legumes. Abiotic Stress and Legumes. Academic Press, US. pp. 93–103.

Bates L S, Waldren R P, Teare I D. 1973. Rapid determination of free proline for water-stress studies. Plant and Soil, 39, 205–207.

Bernfeld P. 1955. Methods in Enzymology. Academic Press, New York. pp. 149–158.

Bradford M M. 1976. A rapid and sensitive method for the quantitation ofmicrogram quantities of protein utilizing the principle of protein-dye binding. Analytical Biochemistry, 72, 248–254.

Chen H N, Tao L Y, Shi J M, Han X R, Cheng X G. 2021. Exogenous salicylic acid signal reveals an osmotic regulatory role in priming the seed germination of Leymus chinensis under salt–alkali stress. Environmental and Experimental Botany, 188, 104498.

Chen K, Arora R. 2013. Priming memory invokes seed stress-tolerance. Environmental and Experimental Botany, 94, 33–45.

Chen Z, Cao X L, Niu J P. 2021. Effects of exogenous ascorbic acid on seed germination and seedling salt-tolerance of alfalfa. PLoS ONE, 16, e0250926.

Cheng B, Li Z, Liang L, Cao Y, Zeng W, Zhang X, Ma X, Huang L, Nie G, Liu W, Peng Y. 2018. The γ-aminobutyric acid (GABA) alleviates salt stress damage during seeds germination of White Clover associated with Na+/K+ transportation, dehydrins accumulation, and stress-related genes expression in White Clover. International Journal of Molecular Sciences, 19, 2520.

Damaris R N, Lin Z, Yang P, He D. 2019. The rice alpha-amylase, conserved regulator of seed maturation and germination. International Journal of Molecular Sciences, 20, 1–17.

Deinlein U, Stephan A B, Horie T, Luo W, Xu G, Schroeder J I. 2014. Plant salt-tolerance mechanisms. Trends in Plant Science, 19, 371–379.

Dubois M, Giles K A, Hamilton J J, Roberes P A, Smith F. 1956. Colorometric method for determination of sugars and related substances. Analytical Chemistry, 28, 350–356.

El-Badri A M, Batool M, Wang C, Hashem A M, Tabl K M, Nishawy E, Kuai J, Zhou G, Wang B. 2021. Selenium and zinc oxide nanoparticles modulate the molecular and morpho-physiological processes during seed germination of Brassica napus under salt stress. Ecotoxicological and Environmental Safety, 225, 112695.

El-Hendawy S, Elshafei A, Al-Suhaibani N, Alotabi M, Hassan W, Dewir Y H, Abdella K. 2019. Assessment of the salt tolerance of wheat genotypes during the germination stage based on germination ability parameters and associated SSR markers. Journal of Plant Interactions, 14, 151–163.

Faizan M, Bhat J A, Chen C, Alyemeni M N, Wijaya L, Ahmad P, Yu F. 2021. Zinc oxide nanoparticles (ZnO-NPs) induce salt tolerance by improving the antioxidant system and photosynthetic machinery in tomato. Plant Physiology and Biochemistry, 161, 122–130.

Fan L Q, Yang L W, Gao H B, Wu X L, Xia Q P, Gong B B. 2012. Effects of exogenous gamma-aminobutyric acid on polyamine metabolism of melon seedlings under hypoxia stress. Journal of Applied Ecology, 23, 1599–1606.

Feghhenabi F, Hadi H, Khodaverdiloo H, van Genuchten M T. 2020. Seed priming alleviated salinity stress during germination and emergence of wheat (Triticum aestivum L.). Agricultural Water Management, 231, 106022.

Gul H, Naz F, Hamayun M, Sayyed A, Sherwani S K. 2014. Effect of NaCl stress on Pisum sativum germination and seedling growth with the influence of seed priming with potassium (KCl and KOH). Eurasian Journal of Agriculture & Environmental Science, 14, 1304–1311.

Guo J, Du M, Tian H, Wang B. 2020. Exposure to high salinity during seed development markedly enhances seedling emergence and fitness of the progeny of the extreme halophyte Suaeda salsa. Frontiers in Plant Science, 11, 1–11.

Hameed A, Sheikh M A, Hameed A, Farooq T, Basra S M A, Jamil A. 2014. Chitosan seed priming improves seed germination and seedling growth in wheat (Triticum aestivum L.) under osmotic stress induced by polyethylene glycol. Philippine Agricultural Scientist, 97, 294–299.

Hemalatha G, Renugadevi J, Evera T. 2017. Seed priming to alleviate the effect of salinity stress in rice. International Journal of Chemical Studies, 5, 1140–1143.

Huang P, He L, Abbas A, Sadam H, Saddam H, Du D, Hafeez M B, Balooch S, Zahra N, Ren X, Rafiq M, Saqib M. 2021. Seed priming with sorghum water extract improves the performance of camelina (Camelina sativa (L.) crantz.) under salt stress. Plants, 10, 749.

Iqbal M, Ashraf M, Jamil A, ur-Rehman S. 2006. Does seed priming induce changes in the levels of some endogenous plant hormones in hexaploid wheat plants under salt stress? Journal of Integrative Plant Biology, 48, 181–189.

Iqbal N, Umar S, Khan N A, Khan M I R. 2014. A new perspective of phytohormones in salinity tolerance: Regulation of proline metabolism. Environmental and Experimental Botany, 100, 34–42.

Jafar M Z, Farooq M, Cheema M A, Afzal I, Basra S M A, Wahid M A, Aziz T, Shahid M. 2012. Improving the performance of wheat by seed priming under saline conditions. Journal of Agronomy and Crop Science, 198, 38–45.

Jini D, Joseph B. 2017. Physiological mechanism of salicylic acid for alleviation of salt stress in rice. Rice Science, 24, 97–108.

Kaneko M, Itoh H, Ueguchi-Tanaka M, Ashikari M, Matsuoka M. 2002. The α-amylase induction in endosperm during rice seed germination is caused by gibberellin synthesized in epithelium. Plant Physiology, 128, 1264–1270.

Khalid M F, Hussain S, Anjum M A, Ejaz S, Ahmad M, Jan M, Zafar S, Zakir I, Ali M A, Ahmad N, Rao M J, Ahmad S. 2019. Hydropriming for plant growth and stress tolerance. In: Hasanuzzaman M, Fotopoulos V, eds., Priming and Pretreatment of Seeds and Seedlings: Implication in Plant Stress Tolerance and Enhancing Productivity in Crop Plants. Springer, Singapore. pp. 373–384.

Khan I, Raza M A, Awan S A, Shah G A, Rizwan M, Ali B, Tariq R, Hassan M J, Alyemeni M N, Brestic M, Zhang X, Ali S, Huang L. 2020. Amelioration of salt induced toxicity in pearl millet by seed priming with silver nanoparticles (AgNPs): The oxidative damage, antioxidant enzymes and ions uptake are major determinants of salt tolerant capacity. Plant Physiology and Biochemistry, 156, 221–232.

Khan M N, Fu C, Li J, Tao Y, Li Y, Hu J, Chen L, Khan Z, Wu H, Li Z. 2023. Seed nanopriming: How do nanomaterials improve seed tolerance to salinity and drought? Chemosphere, 310, 136911.

Khan M N, Li Y, Khan Z, Chen L, Liu J, Hu J, Wu H, Li Z. 2021. Nanoceria seed priming enhanced salt tolerance in rapeseed through modulating ROS homeostasis and α-amylase activities. Journal of Nanobiotechnology, 19, doi: 10.1186/s12951-021-01026-9.

Khan M N, Zhang J, Luo T, Liu J, Rizwan M, Fahad S, Xu Z, Hu L. 2019. Seed priming with melatonin coping drought stress in rapeseed by regulating reactive oxygen species detoxification: Antioxidant defense system, osmotic adjustment, stomatal traits and chloroplast ultrastructure perseveration. Industrial Crops and Products, 140, 111597.

Li Y, Liang L, Li W, Ashraf U, Ma L, Tang X, Pan S, Tian H, Mo Z. 2021. ZnO nanoparticle-based seed priming modulates early growth and enhances physio-biochemical and metabolic profiles of fragrant rice against cadmium toxicity. Journal of Nanobiotechnology, 19, doi: 10.1186/s12951-021-00820-9.

Liu H, Hussain S, Zheng M, Peng S, Huang J, Cui K, Nie L. 2015. Dry seeded rice as an alternative to transplantedflooded rice in Central China. Agronomy for Sustainable Development, 35, 285–294.

Liu J, Li L, Yuan F, Chen M. 2019. Exogenous salicylic acid improves the germination of Limonium bicolor seeds under salt stress. Plant Signal & Behavior, 14, 1–8.

Mahakham W, Sarmah A K, Maensiri S, Theerakulpisut P. 2017. Nanopriming technology for enhancing germination and starch metabolism of aged rice seeds using phytosynthesized silver nanoparticles. Scientific Reports, 7, 1–21.

Mazhar M W, Ishtiaq M, Hussain I, Parveen A, Bhatti K H, Azeem M, Thind S, Ajaib M, Maqbool M, Sardar T, Muzammil K, Nasir N. 2022. Seed nano-priming with zinc oxide nanoparticles in rice mitigates drought and enhances agronomic profile. PLoS ONE, 17, 1–18.

Miladinov Z, Balesevic-Tubic S, Djordjevic V, Djukic V, Ilic A, Cobanovic L. 2015. Optimal time of soybean seed priming and primer effect under salt stress conditions. Journal of Agricultural Science, 60, 109–117.

Miller G, Suzuki N, Ciftci-Yilmaz S, Mittler R. 2010. Reactive oxygen species homeostasis and signalling during drought and salinity stresses. Plant Cell and Environment, 33, 453–467.

Munns R. 2002. Comparative physiology of salt and water stress. Plant Cell and Environment, 25, 239–250.

Munns R, Tester M. 2008. Mechanisms of salinity tolerance. Annual Review of Plant Biology, 59, 651–681.

Mutlu S, Atici Ö, Nalbantoglu B. 2009. Effects of salicylic acid and salinity on apoplastic antioxidant enzymes in two wheat cultivars differing in salt tolerance. Biologi Plantarum, 53, 334–338.

Nasri N, Kaddour R B, Mahmoudi H, Bouraoui N K. 2011. The effect of osmopriming on germination, seedling growth and phosphatase activities of lettuce under saline condition. African Journal of Biotechnology, 10, 14366–14372.

Nciizah A D, Rapetsoa M C, Wakindiki I I, Zerizghy M G. 2020. Micronutrient seed priming improves maize (Zea mays L.) early seedling growth in a micronutrient deficient soil. Heliyon, 6, e04766.

Othman Y, Al-Karaki G, Al-Horani A. 2006. Variation in germination and ion uptake in barley genotypes under salinity conditions. World Journal of Agricultural Sciences, 2, 11–15.

Pál M. 2018. Seed priming, a re-discovered old method. Journal of Soil Plant Biology, 1, 21–23.

Panuccio M R, Chaabani S, Roula R, Muscolo A. 2018. Bio-priming mitigates detrimental effects of salinity on maize improving antioxidant defense and preserving photosynthetic efficiency. Plant Physiology and Biochemistry, 132, 465–474.

Parvaiz A, Satyawati S. 2008. Salt stress and phyto-biochemical responses of plants - A review. Plant Soil and Environment, 54, 89–99.

Pereira A D E S, Oliveira H C, Fraceto L F, Santaella C. 2021. Nanotechnology potential in seed priming for sustainable agriculture. Nanomaterials, 11, 1–29.

Pickson R B, Gui P, Chen A, Boateng E. 2022. Empirical analysis of rice and maize production under climate change in China. Environmental Science Pollution Research, 29, 70242–70261.

Plaksenkova I, Kokina I, Petrova A, Jermaļonoka M, Gerbreders V, Krasovska M. 2020. The impact of zinc oxide nanoparticles on cytotoxicity, genotoxicity, and mirna expression in barley (Hordeum vulgare L.) seedlings. Scientific World Journal, 2020, 6649746.

Pradhan N, Prakash P, Manimurugan C, Tiwari S K, Sharma R P, Singh P. 2015. Screening of tomato genotypes using osmopriming with PEG 6000 under salinity stress. Research Environment Life Sciences, 8, 245–250.

Rehman H U, Iqbal H, Basra S M A, Afzal H, Farooq M, Wakeel A, Wang N. 2015. Seed priming improves early seedling vigor, growth and productivity of spring maize. Journal of Integrative Agriculture, 14, 1745–1754.

Rai-Kalal P, Jajoo A. 2021. Priming with zinc oxide nanoparticles improve germination and photosynthetic performance in wheat. Plant Physiology and Biochemistry, 160, 341–351.

Rajjou L, Debeaujon I. 2008. Seed longevity: Survival and maintenance of high germination ability of dry seeds. Current Biology, 331, 796–805.

Rajjou L, Duval M, Gallardo K, Catusse J, Bally J, Job C, Job D. 2012. Seed germination and vigor. Annual Review Plant Biology, 63, 507–533.

Rico C M, Morales M I, McCreary R, Castillo-Michel H, Barrios A C, Hong J, Tafoya A, Lee W Y, Varela-Ramirez A, Peralta-Videa J R, Gardea-Torresd ey J L. 2013. Cerium oxide nanoparticles modify the antioxidative stress enzyme activities and macromolecule composition in rice seedlings. Environmantal Science & Technology, 47, 14110–14118.

Rossi L, Zhang W, Lombardini L, Ma X. 2016. The impact of cerium oxide nanoparticles on the salt stress responses of Brassica napus L. Environmental Pollution, 219, 28–36.

Sami F, Yusuf M, Faizan M, Faraz A, Hayat S. 2016. Role of sugars under abiotic stress. Plant Physiology and Biochemistry, 109, 54–61.

Sarraf M, Vishwakarma K, Kumar V, Arif N, Das S, Johnson R, Janeeshma E, Puthur J T, Aliniaeifard S, Chauhan D K, Fujita M, Hasanuzzaman M. 2022. Metal/metalloid-based nanomaterials for plant abiotic stress tolerance: An overview of the mechanisms. Plants, 11, 1–31.

Shah T, Latif S, Saeed F, Ali I, Ullah S, Abdullah A A, Jan S, Ahmad P. 2021. Seed priming with titanium dioxide nanoparticles enhances seed vigor, leaf water status, and antioxidant enzyme activities in maize (Zea mays L.) under salinity stress. Journal of King Saud University Science, 33, 101207.

Shereen A, Mumtaz S, Raza S, Khan M A, Solangi S. 2005. Salinity effects on seedling growth and yield components of different inbred rice lines. Pakistan Journal of Botany, 37, 131–139.

Shtull-Trauring E, Cohen A, Ben-Hur M, Tanny J, Bernstein N. 2020. Reducing salinity of treated waste water with large scale desalination. Water Research, 186, 116322.

Szöllösi R, Molnár Á, Kondak S, Kolbert Z. 2020. Dual effect of nanomaterials on germination and seedling growth: Stimulation vs. phytotoxicity. Plants, 9, 1–30.

Wang W, Peng S, Liu H, Tao Y, Huang J, Cui K, Nie L. 2017. The possibility of replacing puddled transplanted flooded rice with dry seeded rice in central China: A review. Field Crops Research, 214, 310–320.

Wang W Q, He A B, Peng S B, Huang J L, Cui K H, Nie L X. 2018. The effect of storage condition and duration on the deterioration of primed rice seeds. Frontiers in Plant Science, 9, 172.

Wang Y, Luo Z, Huang X, Yang K, Gao S, Du R. 2014. Effect of exogenous γ-aminobutyric acid (GABA) treatment on chilling injury and antioxidant capacity in banana peel. Scientia Hortic-Amsterdam, 168, 132–137.

Wang Z, Li H, Li X, Xin C, Si J, Li S, Li Y, Zheng X, Li H, Wei X, Zhang Z, Kong L, Wang F. 2019. Nano-ZnO priming induces salt tolerance by promoting photosynthetic carbon assimilation in wheat. Archives of Agronomy and Soil Science, 66, 1259–1273.

Yang J, Duan G, Li C, Liu L, Han G, Zhang Y, Wang C. 2019. The crosstalks between jasmonic acid and other plant hormone signaling highlight the involvement of jasmonic acid as a core component in plant response to bioticand abiotic stresses. Frontiers in Plant Science, 10, 1–12.

Zheng M, Tao Y, Hussain S, Jiang Q, Peng S, Huang J, Cui K, Nie L. 2016. Seed priming in dry direct-seeded rice: consequences for emergence, seedling growth and associated metabolic events under drought stress. Plant Growth Regulation, 78, 167–178.

Zeng L, Cai J S, Li J J, Lu G Y, Li C S, Fu G P, Zhang X K, Ma H Q, Liu Q Y, Zou X L, Cheng Y. 2018. Exogenous application of a low concentration of melatonin enhances salt tolerance in rapeseed (Brassica napus L.) seedlings. Journal of Integrative Agriculture, 17, 328–335.

Zhang L, Ma H, Chen T, Pen J, Yu S, Zhao X. 2014. Morphological and physiological responses of cotton (Gossypium hirsutum L.) plants to salinity. PLoS ONE, 9, 0112807.

Zhang X, Wang D, Fang F, Zhen Y, Liao X. 2005. Food safety and rice production in China. Research of Agricutural Modernization, 26, 85–88.

Zhao C, Zhang H, Song C, Zhu J K, Shabala S. 2020. Mechanisms of plant responses and adaptation to soil salinity. Innovation (China), 1, 100017.

Zhu J K. 2016. Abiotic stress signaling and responses in plants. Cell, 167, 313–324.