Chu G, Xu R, Chen S, Xu C M, Liu Y H, Zang X F, Wang D Y. 2021. Effects of optimized cultivation patterns on root–shoot growth characteristics, water and nitrogen use efficiency and yield of rice. Chinese Journal of Rice Science, 35, 586–594. (in Chinese)

Deng J, Feng X, Wang D, Lu J, Chong H, Shang C, Liu K, Huang L, Tian X, Zhang Y. 2022. Root morphological traits and distribution in direct-seeded rice under dense planting with reduced nitrogen. PLoS ONE, 15, e0238362.

Doussan C, Pagès L, Pierret A. 2003. Soil exploration and resource acquisition by plant roots: An architectural and modelling point of view. Sustainable Agriculture, 23, 419–431.

Freschet G T, Cyrille V, Bourget M Y, Lorenzen M S, Fort F. 2018. Allocation, morphology, physiology, architecture: The multiple facets of plant above- and below-ground responses to resource stress. New Phytologist, 219, 1142–1144.

Gale M R, Grigal D F. 1987. Vertical root distributions of northern tree species in relation to successional status. Canadian Journal of Forest Research, 17, 829–834.

Guo Q R, Chen L. 2012. Advantages and prospect analysis of rice sub-membrane drip irrigation cultivation technology development in China. China Rice, 18, 36–39. (in Chinese)

Ha S, Vankova R, Yamaguchi-Shinozaki K, Shinozaki K, Tran L S P. 2012. Cytokinins: Metabolism and function in plant adaptation to environmental stresses. Trends in Plant Science, 17, 172–179.

He H B, Ma F Y, Yang R, Chen L, Jia B, Cui J, Fan H, Wang X, Li L. 2013. Rice performance and water use efficiency under plastic mulching with drip irrigation. PLoS ONE, 8, e83103.

Ji L, Li Y X, Zhang X Z, Yu H Y. 2012. Root morphology and activity characteristics of nitrogen efficient utilization genotype rice. Scientia Agricultura Sinica, 45, 4770–4781. (in Chinese)

Ju X T, Gu B J. 2014. Current situation, problems and trends of nitrogen fertiliser application on farmland in China. Journal of Plant Nutrition and Fertilizers, 20, 783–795. (in Chinese)

Kramer-Walter K R, Bellingham P J, Millar T R, Smissen R D, Richardson S J, Laughlin D C. 2016. Root traits are multidimensional: Specific root length is independent from root tissue density and the plant economic spectrum. Journal of Ecology, 104, 1299–1310.

Li H N, Xu H G, Ren Y H, Ding N, Jiang Y M. 2015. Effects of different nitrogen application levels on growth, nitrogen utilization and endogenous hormone content of dwarf Fu-ji apple saplings. Journal of Plant Nutrition and Fertilizer, 21, 1304–1311. (in Chinese)

Li N, Yang Z Y, Dai Z, Sun Y J, Xu H, He Y, Yan T R, Jiang M J, Guo C C, Wang C Y, Ma J. 2017. Relationship between root morphology, nitrogen uptake and utilization and yield of rice with different nitrogen efficiency. Scientia Agricultura Sinica, 50, 2683–2695. (in Chinese)

Liu K, Chen Y, Li S Y, Wang W L, Zhang W Y, Zhang H. 2023. Differing responses of root morphology and physiology to nitrogen application rates and their relationships with grain yield in rice. The Crop Journal, 11, 618–627.

Liu K, Li T T, Chen Y, Huang J, Qiu Y Y, Li S Y, Wang H, Zhu A, Zhou X X, Yu F, Zhang H, Gu J F, Liu L J, Yang J C. 2020. Effects of root morphology and physiology on the formation and regulation of large panicles in rice. Field Crops Research, 258, 107–946.

Milla R, Osborne C P, Turcotte M M, Violle C. 2015. Plant domestication through an ecological lens. Trends in Ecology & Evolution, 30, 463–469.

Mofijul Islam S M, Gaihre Y K, Biswas J C, Jahan M S, Singh U, Adhikary S K, Satter M A, Saleque M A. 2018. Different nitrogen rates and methods of application for dry season rice cultivation with alternate wetting and drying irrigation: Fate of nitrogen and grain yield. Agricultural Water Management, 196, 144–153.

National Bureau of Statistics of China. 2022. China Statistical Yearbook. https://www.stats.gov.cn/gk/ (in Chinese)

Pandey V, Shukla A. 2015. Acclimation and tolerance strategies of rice under drought stress. Rice Science, 22, 147–161.

Roucou A, Violle C, Fort F, Roumet P, Ecarnot M, Vile D. 2018. Shifts in plant functional strategies over the course of wheat domestication. Journal of Applied Ecology, 55, 25–37.

Takei K, Takahashi T, Sugiyama T, Yamaya T, Sakakibara H. 2002. Multiple routes communicating nitrogen availability from roots to shoots: A signal transduction pathway mediated by cytokinin. Journal of Experimental Botany, 53, 971–977.

Talla S K, Panigrahy M, Kappara S, Nirosha P, Neelamraju S, Ramanan R. 2016. Cytokinin delays dark-induced senescence in rice by maintaining the chlorophyll cycle and photosynthetic complexes. Journal of Experimental Botany, 67, 1839–1851.

Walch-Liu P, Ivanov I, Filleur S Y, Gan Y, Remans T, Forde B G. 2006. Nitrogen regulation of root branching. Annals of Botany, 97, 875–881.

Wang X X, Li H B, Chu Q, Feng G, Kuyper T W, Rengel Z. 2020. Mycorrhizal impacts on root trait plasticity of six maize cultivars along a phosphorus supply gradient. Plant and Soil, 448, 71–86.

Wen Z H, Li H B, Shen Q, Tang X M, Xiong C Y, Li H G, Pang J Y, Ryan M H, Lambers H, Shen J B. 2019. Tradeoffs among root morphology, exudation and mycorrhizal symbioses for phosphorus-acquisition strategies of 16 crop species. New Phytologist, 223, 882–895.

Wen Z H, Pang J Y, Tueux G, Liu Y F, Shen J B, Ryan M H, Lambers H, Siddique K H M. 2020. Contrasting patterns in biomass allocation, root morphology and mycorrhizal symbiosis for phosphorus acquisition among 20 chickpea genotypes with different amounts of rhizosheath carboxylates. Functional Ecology, 34, 1311–1324.

Xu G, Jiang M M, Lu D K, Wang H Z, Chen M C. 2020. Nitrogen forms affect the root characteristic, photosynthesis, grain yield, and nitrogen use efficiency of rice under different irrigation regimes. Crop Science, 60, 2594–2610.

Xu G W, Lu D K, Wang H Z. 2017. Coupling effect of wetting and drying alternative irrigation and nitrogen application rate on photosynthetic characteristics of rice leaves. Plant Nutrition and Fertilizer Science, 23, 1225–1237. (in Chinese)

Xu G W, Lu D K, Wang H Z, Li Y J. 2018. Morphological and physiological traits of rice roots and their relationships to yield and nitrogen utilization as influenced by irrigation regime and nitrogen rate. Agriculture Water Manage, 203, 385–394.

Yang J C, Zhang H, Zhang J H. 2012. Root morphology and physiology in relation to the yield formation of rice. Journal of Integrative Agriculture, 11, 920–926.

Yang J C, Zhou Q, Zhang J H. 2017. Moderate wetting and drying increases rice yield and reduces water use, grain arsenic level, and methane emission. The Crop Journal, 5, 151–158.

Zhang S W, He Q L, Wang H Y, Jiang M J, Li Y H, Sun Y J, Guo X, Ma J. 2018. Effects of nitrogen application rates on nitrogen use efficiency and grain yield of indica hybrid rice F You498 under controlled intermittent irrigation. Journal of Plant Nutrition and Fertilizers, 24, 82–94. (in Chinese)

Zhang W Y, Yu J X, Xu Y J, Wang Z Q, Liu L J, Zhang H, Gu J F, Zhang J H, Yang J C. 2021. Alternate wetting and drying irrigation combined with the proportion of polymer-coated urea and conventional urea rates increases grain yield, water and nitrogen use efficiencies in rice. Field Crops Research, 265, 108–165.

Zhang Y J, Xu J N, Cheng Y D, Wang C, Liu G S, Yang J C. 2020. The effects of water and nitrogenon the roots and yield of upland and paddy rice. Journal of Integrative Agriculture, 19, 1363–1374.

Zhou Y F, Wang D Q, Lu Z B, Wang N, Wang Y T, Li F X, Xu W J, Huang R D. 2014. Effects of drought stress on photosynthetic characteristics and endogenous hormone ABA and CTK contents in green-stayed sorghum. Scientia Agricultura Sinica, 47, 655–663. (in Chinese)

Zörb C, Ludewig U, Hawkesford M J. 2018. Perspective on wheat yield and quality with reduced nitrogen supply. Trends in Plant Science, 23, 1029–1037.