Ali Y, Atta B M, Akhter J, Monneveux P, Lateef Z. 2008. Genetic variability, association and diversity studies in wheat (Triticum aestivum L.) germplasm. Pakistan Journal of Botany, 40, 2087–2097.

Ariel F, Roxana S, Slafer G A. 2013. Floret development and grain setting differences between modern durum wheats under contrasting nitrogen availability. Journal of Experimental Botany, 64, 169–184.

Arnon D I. 1949. Copper enzymes in isolated chloroplasts. Polyphenoloxidase in Beta vulgaris. Plant Physiology, 24, 1–15.

Borràs-Gelonch G, Rebetzke G J, Richards R A, Romagosa I. 2012. Genetic control of duration of pre-anthesis phases in wheat (Triticum aestivum L.) and relationships to leaf appearance, tillering, and dry matter accumulation. Journal of Experimental Botany, 63, 69–89.

Brinton J, Uauy C. 2019. A reductionist approach to dissecting grain weight and yield in wheat. Journal of Integrative Plant Biology, 61, 337–358.

Carmo-Silva E, Andralojc P J, Scales J C, Driever S M, Mead A, Lawson T, Raines C A, Parry M A J. 2017. Phenotyping of field-grown wheat in the UK highlights contribution of light response of photosynthesis and flag leaf longevity to grain yield. Journal of Experimental Botany, 68, 3473–3486.

Chan K Y, Bowman A, Oates A. 2001. Oxidizible organic carbon fractions and soil quality changes in an oxic paleustalf under different pasture leys. Soil Science, 166, 61–67.

Chen J, Fei K Q, Zhang W Y, Wang Z L, Zhang J H, Yang J C. 2021. Brassinosteroids mediate the effect of high temperature during anthesis on the pistil activity of photo-thermosensitive genetic male-sterile rice lines. The Crop Journal, 9, 109–119.

Dhyani K, Ansari M W, Rao Y R, Verma R S, Shukla A, Tuteja N. 2013. Comparative physiological response of wheat genotypes under terminal heat stress. Plant Signaling & Bbehavior, 8, e24564.

Dreccer M F, Wockner K B, Palta J A, McIntyre C L, Borgognone M G, Bourgault M, Reynolds M, Miralles D J. 2014. More fertile florets and grains per spike can be achieved at higher temperature in wheat lines with high spike biomass and sugar content at booting. Functional Plant Biology, 41, 482–495.

Duan J, Wu Y, Zhou Y, Ren X, Shao Y, Feng W, Zhu Y, Wang Y, Guo T C. 2018. Grain number responses to pre-anthesis dry matter and nitrogen in improving wheat yield in the Huang-Huai Plain. Scientific Reports, 8, 7126.

Evans J R, Clarke V C. 2019. The nitrogen cost of photosynthesis. Journal of Experimental Botany, 70, 7–15.

Ferrante A, Savin R, Slafer G A. 2013. Is floret primordia death triggered by floret development in durum wheat? Journal of Experimental Botany, 64, 2859–2869.

Fischer R A. 2008. The importance of grain or kernel number in wheat: A reply to Sinclair and Jamieson. Field Crops and Research, 105, 15–21.

Foulkes M J, Slafer G A, Davies W J, Berry P M, Sylvester-Bradley R, Martre P, Calderini D F, Griffiths S, Reynlods M P. 2011. Raising yield potential of wheat. III. Optimizing partitioning to grain while maintaining lodging resistance. Journal of Experimental Botany, 62, 469–486.

Gautrat P, Laffont C, Frugier F, Ruffel S. 2021. Nitrogen systemic signaling: From symbiotic nodulation to root acquisition. Trends in Plant Science, 26, 392–406.

Ghiglione H O, Gonzalez F G, Serrago R, Maldonado S B, Chilcott C, Cura J A, Miralles D J, Zhu T, Casal J J. 2008. Autophagy regulated by day length determines the number of fertile florets in wheat. The Plant Journal, 55, 1010–1024.

Grote U, Fasse A, Nguyen T T, Erenstein O. 2021. Food security and the dynamics of wheat and maize value chains in Africa and Asia. Frontiers in Sustainable Food Systems, 4, 617009.

Han B, Fu L, Zhang D, He X, Chen Q, Peng M, Zhang J. 2016. Interspecies and intraspecies analysis of trehalose contents and the biosynthesis pathway gene family reveals crucial roles of trehalose in osmotic-stress tolerance in Cassava. International Journal of Molecular Sciences, 17, 1077.

Hawkesford M J. 2014. Reducing the reliance on nitrogen fertilizer for wheat production. Journal of Cereal Science, 59, 276–283.

Hou L, Lin R X, Wang X J, Li H, Zhao C Z, Zhu X J, Li C S, Li G H. 2022. The mechanisms of pod zone nitrogen application on peanut pod yield. Russian Journal of Plant Physiology, 69, 51.

Hütsch B W, Jahn D, Schubert S. 2019. Grain yield of wheat (Triticum aestivum L.) under long-term heat stress is sink-limited with stronger inhibition of kernel setting than grain filling. Journal of Agronomy & Crop Science, 205, 22–32.

Jiang D, Fan X, Dai T, Cao W. 2008. Nitrogen fertiliser rate and post-anthesis waterlogging effects on carbohydrate and nitrogen dynamics in wheat. Plant and Soil, 304, 301–314.

Ju X T, Xing G X, Chen X P, Zhang L J, Liu X J, Cui Z L, Yin B, Christie P, Zhu Z L, Zhang F S. 2009. Reducing environmental risk by improving N management in intensive Chinese agricultural systems. Proceedings of the National Academy of Sciences of the United States of America, 106, 3041–3046.

Kuang T Y, Peng D C, Tang C Q, Li T Z, Lou S Q, Zhang Q D, Zuo B Y, Lin S Q. 1990. The organization of chlorophyll-proteins of PSII and PSI on thylakoid membrane. Current Research in Photosynthesis, 2, 335–338.

Liang W, Zhang Z, Wen X, Liao Y, Liu Y. 2017. Effect of non-structural carbohydrate accumulation in the stem pre-anthesis on grain filling of wheat inferior grain. Field Crops Research, 211, 66–76.

Liu Y, Wang Q, Ding Y, Li G, Xu J, Wang S. 2011. Effects of external ABA, GA3 and NAA on the tiller bud outgrowth of rice is related to changes in endogenous hormones. Plant Growth Regulation, 65, 247–254.

Lisso J, Altmann T, Müssig C. 2006. Metabolic changes in fruits of the tomato dx mutant. Phytochemistry, 6720, 2232–2238.

Lopes M S, Cortadellas N, Kichey T, Dubois F, Habash D Z, Araus J L. 2006. Wheat nitrogen metabolism during grain filling: Comparative role of glumes and the flag leaf. Planta, 225, 165–181.

Lu D, Lu F, Pan J, Cui Z, Zou C, Chen X, He M, Wang Z. 2015. The effects of cultivar and nitrogen management on wheat yield and nitrogen use efficiency in the North China Plain. Field Crops Research, 171, 157–164.

Maria S D, Puschenreiter M, Puschenreiter A R. 2013. Cadmium accumulation and physiological response of sunflower plants to Cd during the vegetative growing cycle. Plant, Soil and Environment (Czech Republic), 59, 254–261.

McIntyre G I. 2001. Control of plant development by limiting factors: A nutritional perspective. Physiologia Plantarum, 113, 165–175.

Noor H, Sun M, Algwaiz H I M, Sher A, Fiaz S, Attia K A, Wani S H, Alkahtani M D F, Husnain L A, Lin W, Gao Z Q. 2022. Chlorophyll fluorescence and grain filling characteristic of wheat (Triticum aestivum L.) in response to nitrogen application level. Molecular Biology Reports, 49, 7157–7172.

Nuttall J G, Barlow K M, Delahunty A J, Christy B P, O’Leary G J. 2018. Acute high temperature response in wheat. Agronomy Journal, 110, 1296–1308.

Pociecha E, Jurczyk B, Dziurka M, Paczyński R, Okleštková J, Janeczko A. 2015. 24-Epibrassinolide promotes carbohydrates accumulation in crowns of perennial ryegrass during cold acclimation by regulation of gene expression and enzyme activities which results in increased frost tolerance. Procedia Environmental Sciences, 29, 234–235.

Saville R J, Gosman N, Burt C J, Makepeace J, Steed A, Corbitt M, Chandler E, Brown J K M, Boulton M I, Nicholson P. 2012. The ‘Green Revolution’ dwarfing genes play a role in disease resistance in Triticum aestivum and Hordeum vulgare. Journal of Experimental Botany, 63, 1271–1283.

Schilling G, Schiller C, Otto S. 1991. Influence of brassinosteroids on organ relations and enzyme activities of sugar-beet plants. Biology, 74, 208–219.

Serrago R A, Miralles D J, Slafer G A. 2008. Floret fertility in wheat as affected by photoperiod during stem elongation and removal of spikelets at booting. European Journal of Agronomy, 28, 301–308.

Steinfort U, Fukai S, Trevaskis B, Glassop D, Chan A, Dreccer M F. 2017. Vernalisation and photoperiod sensitivity in wheat: The response of floret fertility and grain number is affected by vernalisation status. Field Crops and Research, 203, 243–255.

Tanz S K, Kilian J, Johnsson C, Apel K, Small I, Harter K, Wanke D, Pogson B, Albrecht V. 2012. The SCO2 protein disulphide isomerase is required for thylakoid biogenesis and interacts with LCHB1 chlorophyll a/b binding proteins which affects chlorophyll biosynthesis in Arabidopsis seedlings. The Plant Journal, 69, 743–754.

Waddington S R, Cartwright P M , Wall P C. 1983. A quantitative scale of spike initial and pistil development in barley and wheat. Annals of Botany, 51, 119–130.

Wang H, Zhang Y, Chen A, Liu H, Zhai L, Lei B, Ren T. 2017. An optimal regional nitrogen application threshold for wheat in the North China Plain considering yield and environmental effects. Field Crops Research, 207, 52–61.

Wang M, Jiang W, Yu H. 2010. Effects of exogenous epibrassinolide on photosynthetic characteristics in tomato (Lycopersicon esculentum Mill) seedlings under weak light stress. Journal of Agricultural and Food Chemistry, 58, 3642–3645.

Wim V D E, Joke D R, André V L. 1999. Effect of nitrogen concentration on fructan and fructan metabolizing enzymes in young chicory plants (Cichorium intybus). Physiologia Plantarum, 105, 2–8.

Wu X, Miao L, Li C S, Mchugh A D, Li M, Xing T, Liu Y B, Tang Y L. 2022. Source–sink relations and responses to sink–source manipulations during grain filling in wheat. Journal of Integrative Agriculture, 21, 1593–1605.

Xia X J, Huang L F, Zhou Y H, Mao W H, Shi K, Wu J X, Asami T, Chen Z X, Yu J Q. 2009a. Brassinosteroids promote photosynthesis and growth by enhancing activation of Rubisco and expression of photosynthetic genes in Cucumis sativus. Planta, 230, 1185–1196.

Xia X J, Wang Y J, Zhou Y H, Tan Y, Mao W H, Shi K, Asami T, Chen Z X, Yu J Q. 2009b. Reactive oxygen species are involved in brassinosteroid-induced stress tolerance in cucumber. Plant Physiology, 150, 801–814.

Xu F, Xi Z, Zhang H, Zhang C, Zhang Z. 2015. Brassinosteroids are involved in controlling sugar unloading in Vitis vinifera ‘Cabernet Sauvignon’ berries during véraison. Plant Physiology and Biochemistry, 94, 197–208.

Yang D, Cai T, Luo Y, Wang Z. 2019. Optimizing plant density and nitrogen application to manipulate tiller growth and increase grain yield and nitrogen-use efficiency in winter wheat. PeerJ, 7, e6484.

Zahedipour-Sheshglani P, Asghari M. 2020. Impact of foliar spray with 24-epibrassinolide on yield, quality, ripening physiology and productivity of the strawberry. Scientia Horticulturae, 268, 109376.

Zhang J H, Liu J L, Zhang J B, Zhao F T, Wang W P. 2010. Effects of nitrogen application rates on translocation of dry matter and nitrogen utilization in rice and wheat. Acta Agronomica Sinica, 36, 1736–1742. (in Chinese)

Zhang W Y, Fu L D, Men C B, Yu J X, Yao J Y, Sheng J Y, Xu Y J, Wang Z Q, Liu L J, Yang J C. 2020a. Response of brassinosteroids to nitrogen rates and their regulation on rice spikelet degeneration during meiosis. Food and Energy Security, 9, e201.

Zhang W Y, Huang H H, Zhou Y J, Zhu K Y, Wu Y F, Xu Y J, Wang W L, Zhang H, Gu J K, Xiong F, Wang Z Q, Liu L J, Yang J C. 2023. Brassinosteroids mediate moderate soil-drying to alleviate spikelet degeneration under high temperature during meiosis of rice. Plant, Cell & Environment, 46, 1340–1362.

Zhang W Y, Sheng J Y, Fu L D, Xu Y J, Xiong F, Wu Y F, Wang W L, Wang Z Q, Zhang J H, Yang J C. 2020b. Brassinosteroids mediate the effect of soil-drying during meiosis on spikelet degeneration in rice. Environmental and Experimental Botany, 169, 103887.

Zhang Z, Huang J, Gao Y, Liu Y, Li J, Zhou X, Yao C, Wang Z, Sun Z, Zhang Y. 2020. Suppressed ABA signal transduction in the spike promotes sucrose use in the stem and reduces grain number in wheat under water stress. Journal Experimental Botany, 71, 7241–7256.

Zhen S M, Deng X, Zhang M, Zhu G R, Lv D W, Wang Y P, Zhu D, Yan Y M. 2017. Comparative phosphoproteomic analysis under high-nitrogen fertilizer reveals central phosphoproteins promoting wheat grain starch and protein synthesis. Frontiers in Plant Science, 8, 67.

Zheng C F, Zhu Y J, Wang C Y, Guo T C. 2016. Wheat grain yield increase in response to pre-anthesis foliar application of 6-benzylaminopurine is dependent on floret development. PLoS ONE, 11, e0156627.

Zheng M J, Chen J, Shi Y H, Li Y X, Yin Y P, Yang D Q, Luo Y L, Pang D W, Xu X, Li W Q, Ni J, Wang Y Y, Wang Z L, Li Y. 2017. Manipulation of lignin metabolism by plant densities and its relationship with lodging resistance in wheat. Scientific Reports, 7, 41805.

Zhou H L, Yao, X D, Zhao Q, Zhang W, Zhang B, Xie F T. 2019. Rapid effect of nitrogen supply for soybean at the beginning flowering stage on biomass and sucrose metabolism. Scientific Reports, 9, 15530.