Bicego B, Savin R, Girousse C, Allard V, Slafer G A. 2024. Tillering and floret development dynamics in wheat cultivars of contrasting spike fertility plasticity. Field Crops Research, 319, 109654.

Cao Y, Zhong Z, Wang H, Shen R. 2022. Leaf angle: A target of genetic improvement in cereal crops tailored for high‐density planting. Plant Biotechnology Journal, 20, 426–436.

Devireddy A R, Zandalinas S I, Gómez-Cadenas A, Blumwald E, Mittler R. 2018. Coordinating the overall stomatal response of plants: Rapid leaf-to-leaf communication during light stress. Science Signaling, 11, eaam9514.

Earle J, Salvatore L, van den Herik B, ten Tusscher K, van Veen H. 2026. How coupling resources and development ensures whole-plant homeostasis. Journal of Experimental Botany, 77, 1452-1468.

Fernández-Milmanda G L, Ballaré C L. 2021. Shade avoidance: Expanding the color and hormone palette. Trends in Plant Science, 26, 509–523.

Frak E, Le Roux X, Millard P, Adam B, Dreyer E, Escuit C, Sinoquet H, Vandame M, Varlet-Grancher C. 2002. Spatial distribution of leaf nitrogen and photosynthetic capacity within the foliage of individual trees: Disentangling the effects of local light quality, leaf irradiance, and transpiration. Journal of Experimental Botany, 53, 2207–2216.

Guo X, Zhang Z, Li J, Zhang S, Sun W, Xiao X, Sun Z, Xue X, Wang Z, Zhang Y. 2024. Phenotypic and transcriptome profiling of spikes reveals the regulation of light regimens on spike growth and fertile floret number in wheat. Plant, Cell & Environment, 47, 1575–1591.

Hao C, Zhu L, Li X, Wang L, Song J, Zhao X, Qin X. 2025. Utilising far‐red light: Photosynthetic and physiological adaptations in shade‐tolerant Fittonia albivenis. Plant, Cell & Environment, 48, 8161–8176.

Holalu S V, Finlayson S A. 2017. The ratio of red light to far red light alters Arabidopsis axillary bud growth and abscisic acid signalling before stem auxin changes. Journal of Experimental Botany, 68, 943–952.

Hu C, Nawrocki W J, Croce R. 2021. Long-term adaptation of Arabidopsis thaliana to far-red light. Plant, Cell & Environment, 44, 3002–3014.

Hu J, Ren B, Chen Y, Liu P, Zhao B, Zhang J. 2022. Exogenous 6-benzyladenine improved the ear differentiation of waterlogged summer maize by regulating the metabolism of hormone and sugar. Frontiers in Plant Science, 13, 848989.

Huber S C, Israel D W. 1982. Biochemical basis for partitioning of photosynthetically fixed carbon between starch and sucrose in soybean (Glycine max Merr.) leaves. Plant Physiology, 69, 691–696.

Izydorczyk C, Nguyen T N, Jo S, Son S, Tuan P A, Ayele B T. 2018. Spatiotemporal modulation of abscisic acid and gibberellin metabolism and signalling mediates the effects of suboptimal and supraoptimal temperatures on seed germination in wheat (Triticum aestivum L.). Plant, Cell & Environment, 41, 1022–1037.

Javid M G, Sorooshzadeh A, Sanavy S A M M, Allahdadi I, Moradi F. 2011. Effects of the exogenous application of auxin and cytokinin on carbohydrate accumulation in grains of rice under salt stress. Plant Growth Regulation, 65, 305–313.

Kabir M R, Nonhebel H M, Backhouse D, Winter G. 2021. Expression of key auxin biosynthesis genes correlates with auxin and starch content of developing wheat (Triticum aestivum L.) grains. Functional Plant Biology, 48, 802–814.

Kebrom T H, Brutnell T P. 2007. The molecular analysis of the shade avoidance syndrome in the grasses has begun. Journal of Experimental Botany, 58, 3079–3089.

Küpers J J, Oskam L, Pierik R. 2020. Photoreceptors regulate plant developmental plasticity through auxin. Plants, 9, 940.

Lei K, Hu H, Chang M, Sun C, Ullah A, Yu J, Dong C, Gao Q, Jiang D, Cao W, Tian Z, Dai T. 2024. A low red/far-red ratio restricts nitrogen assimilation by inhibiting nitrate reductase associated with downregulated TaNR1.2 and upregulated TaPIL5 in wheat (Triticum aestivum L.). Plant Physiology and Biochemistry, 206, 107850.

Lei K, Tan Q, Zhu L, Xu L, Yang S, Hu J, Gao L, Hou P, Shao Y, Jiang D, Cao W, Dai T, Tian Z. 2022. Low red/far-red ratio can induce cytokinin degradation resulting in the inhibition of tillering in wheat (Triticum aestivum L.). Frontiers in Plant Science, 13, 971003.

Leivar P, Monte E. 2014. PIFs: Systems integrators in plant development. The Plant Cell, 26, 56–78.

Li J, Liang Z, Li Y, Wang K, Li Y, Nangia V, Mo F, Liu Y. 2024. Advantageous spike-to-stem competition for assimilates contributes to the reduction in grain number loss in wheat spikes under water deficit stress. Agricultural Water Management, 292, 108675. 

Li L, Ljung K, Breton G, Schmitz R J, Pruneda-Paz J, Cowing-Zitron C, Cole B J, Ivans L J, Pedmale U V, Jung H S, Ecker J R, Kay S A, Chory J. 2012. Linking photoreceptor excitation to changes in plant architecture. Genes & Development, 26, 785–790.

Li X, Zhang X, Bie X, Zhang J, Jiang D J, Tang H, Wang F. 2024. Transcriptome analysis of axillary buds in low phosphorus stress and functional analysis of TaWRKY74s in wheat. BMC Plant Biology, 24, 1.

Liang Z, Cao X, Gao R, Guo N, Tang Y, Nangia V, Liu Y. 2025. Brassinosteroids alleviate wheat floret degeneration under low nitrogen stress by promoting the redistribution of sucrose from stems to spikes. Journal of Integrative Agriculture, 24, 497–516.

Liu Q, Guo Y, Yin P, Lan T, Wei G, Liu F, Lyu C, Kong F, Yuan J. 2025. Low-nitrogen-tolerant maize cultivar improves photosynthesis and yield by optimizing nitrogen allocation to increase electron transport under low-nitrogen and low-light stress. The Crop Journal, 13, 1607–1618. 

Liu T, Hu Y, Li X. 2008. Comparison of dynamic changes in endogenous hormones and sugars between abnormal and normal Castanea mollissima. Progress in Natural Science, 18, 685–690.

Liu Y, Gu D, Ding Y, Wang Q, Li G, Wang S. 2011. The relationship between nitrogen, auxin and cytokinin in the growth regulation of rice (Oryza sativa L.) tiller buds. Australian Journal of Crop Science, 5, 1019-1026.

Liu Y, Zhang L, Zhu A, Shen L, Zhang J, Chen J, Chang G, Yin C, Wang Z, Sun Z, Shen K, Xu X, Sun M, Xin M, Wu J, Lu Z, Tong Y, He Z, Lu F, Hao Y, et al. 2025. Multi-omics identifies key genetic and metabolic networks regulating spike organ development in wheat. The Plant Cell, 37, koaf250.

Liu Z, Hu C, Wang Y, Liu Z, Sha Y, Hao Z, Chen F, Yuan L, Mi G. 2021. Nitrogen allocation and remobilization contributing to low-nitrogen tolerance in stay-green maize. Field Crops Research, 263, 108078.

Livak K J, Schmittgen T D. 2001. Analysis of relative gene expression data using real-time quantitative PCR and the 2^−ΔΔCT method. Methods, 25, 402–408.

Love B, Molero G, Rivera-Amado C, Müller M, Munné-Bosch S, Reynolds M P, Foulkes M J. 2024. Associations between endogenous spike cytokinins and grain-number traits in spring wheat genotypes. European Journal of Agronomy, 152, 127011.

Luo X, Yang Y, Lin X, Xiao J. 2023. Deciphering spike architecture formation towards yield improvement in wheat. Journal of Genetics and Genomics, 50, 835–845.

Pan X, Welti R, Wang X. 2010. Quantitative analysis of major plant hormones in crude plant extracts by high-performance liquid chromatography-mass spectrometry. Nature Protocols, 5, 986–992.

Paolacci A R, Tanzarella O A, Porceddu E, Ciaffi M. 2009. Identification and validation of reference genes for quantitative RT-PCR normalization in wheat. BMC Molecular Biology, 10, 11.

Pérez-Alfocea F, Albacete A, Ghanem M E, Dodd I C. 2010. Hormonal regulation of source-sink relations to maintain crop productivity under salinity: A case study of root-to-shoot signalling in tomato. Functional Plant Biology, 37, 592–603.

Postma J A, Hecht V L, Hikosaka K, Nord E A, Pons T L, Poorter H. 2021. Dividing the pie: A quantitative review on plant density responses. Plant, Cell & Environment, 44, 1072–1094.

Reddy S K, Holalu S V, Casal J J, Finlayson S A. 2013. Abscisic acid regulates axillary bud outgrowth responses to the ratio of red to far-red light. Plant Physiology, 163, 1047–1058.

Schaller G E, Street I H, Kieber J J. 2014. Cytokinin and the cell cycle. Current Opinion in Plant Biology, 21, 7–15.

Shang Y, Wang S, Lin X, Gu S, Wang D. 2023. Supplemental irrigation at jointing improves spike formation of wheat tillers by regulating sugar distribution in ear and stem. Agricultural Water Management, 279, 108160.

Shao A, Ma W, Zhao X, Hu M, He X, Teng W, Li H, Tong Y. 2017. The auxin biosynthetic TRYPTOPHAN AMINOTRANSFERASE RELATED TaTAR2.1-3A increases grain yield of wheat. Plant Physiology, 174, 2274–2288.

Son S, Chitnis V R, Liu A, Gao F, Nguyen T N, Ayele B T. 2016. Abscisic acid metabolic genes of wheat (Triticum aestivum L.): Identification and insights into their functionality in seed dormancy and dehydration tolerance. Planta, 244, 429–447.

Song X, Gu X, Chen S, Qi Z, Yu J, Zhou Y, Xia X. 2024. Far‐red light inhibits lateral bud growth mainly through enhancing apical dominance independently of strigolactone synthesis in tomato. Plant, Cell & Environment, 47, 429–441.

Sparkes D L, Holme S J, Gaju O. 2006. Does light quality initiate tiller death in wheat? European Journal of Agronomy, 24, 212–217.

Tilley M S, Heiniger R W, Crozier C R. 2019. Tiller initiation and its effects on yield and yield components in winter wheat. Agronomy Journal, 111, 1323–1332.

Ugarte C C, Trupkin S A, Ghiglione H, Slafer G, Casal J J. 2010. Low red/far-red ratios delay spike and stem growth in wheat. Journal of Experimental Botany, 61, 3151–3162.

Wang Z, Cao W, Dai T, Zhou Q. 2001. Effects of exogenous hormones on floret development and grain setin wheat. Plant Growth Regulation, 35, 225–231.

Wassenaar M L J, Van Ieperen W, Driever S M. 2022. Low red to far-red ratio increases resistance to CO₂ diffusion and reduces photosynthetic efficiency in low light grown tomato plants. Environmental and Experimental Botany, 200, 104918.

Wen L, Liu Y, Zhou B, Sun W, Xiao X, Wang Z, Sun Z, Zhang Z, Zhang Y. 2024. Differences between two wheat genotypes in the development of floret primordia and contents of pigments and hormones. The Crop Journal, 12, 1196–1207.

Xu H, Cai T, Wang Z, He M. 2015. Physiological basis for the differences of productive capacity among tillers in winter wheat. Journal of Integrative Agriculture, 14, 1958–1970.

Yang H, Li Y, Qiao Y, Sun H, Liu W, Qiao W, Li W, Liu M, Dong B. 2023. Low light stress promotes new tiller regeneration by changing source-sink relationship and activating expression of expansin genes in wheat. Plant, Cell & Environment, 46, 1562–1581.

Yao C, Finlayson S A. 2015. Abscisic acid is a general negative regulator of Arabidopsis axillary bud growth. Plant Physiology, 169, 611–626.

Yao Z, Shi Y, Zou Y, Meng L. 2025. The interaction between ABA and sugar signalling regulates stomatal production in systemic leaves by controlling sucrose transport. Plant, Cell & Environment, 48, 3902–3912.

Youssef H M, Hansson M. 2019. Crosstalk among hormones in barley spike contributes to the yield. Plant Cell Reports, 38, 1013–1016.

Zhang N, Liu Y, Gui S, Wang Y. 2025. Regulation of tillering and panicle branching in rice and wheat. Journal of Genetics and Genomics, 52, 869–886.

Zhang W, Peng K, Cui F, Wang D, Zhao J, Zhang Y, Yu N, Wang Y, Zeng D, Wang Y, Cheng Z, Zhang K. 2021. Cytokinin oxidase/dehydrogenase OsCKX11 coordinates source and sink relationship in rice by simultaneous regulation of leaf senescence and grain number. Plant Biotechnology Journal, 19, 335–350.

Zhang Z, Huang J, Gao Y, Liu Y, Li J, Zhou X, Yao C, Wang Z, Sun Z, Zhang Y. 2020. Suppressed ABA signal transduction in the spike promotes sucrose use in the stem and reduces grain number in wheat under water stress. Journal of Experimental Botany, 71, 7241–7256.