Arif M, Ilyas M, Riaz M, Ali K, Shah K, Haq I U, Fahad S. 2017. Biochar improves phosphorus use efficiency of organic-inorganic fertilizers, maize-wheat productivity and soil quality in a low fertility alkaline soil. Field Crops Research, 214, 25-37.

Cao N, Chen G D, Wang S, Li H Q, Lin J, Hu Q, Wan S M. 2025. Optimizing plant density and nitrogen fertilization in jujube/cotton intercropping systems for sustainable yield and reduced greenhouse gas emissions. Field Crops Research, 326, 109873.

Chapagain T, Riseman A. 2015. Nitrogen and carbon transformations, water use efficiency and ecosystem productivity in monocultures and wheat-bean intercropping systems. Nutrient Cycling in Agroecosystems, 101, 107-121.

Chapepa B, Mudada N, Mapuranga R. 2020. The impact of plant density and spatial arrangement on light interception on cotton crop and seed cotton yield: An overview. Journal of Cotton Research, 3, 18.

Chen H, Wu W, Du K, Yang J, Kang X. 2025. CCT39 transcription factor promotes chlorophyll biosynthesis and photosynthesis in poplar. Plant, Cell & Environment, 48, 3136-3150.

Cheng B, Wang L, Liu R J, Wang W B, Yu R W, Zhou T, Ahmad I, Raza A, Jiang S J, Xu M, Liu C Y, Yu L, Wang W Y, Jing S Z, Liu W G, Yang W Y. 2022. Shade-tolerant soybean reduces yield loss by regulating its canopy structure and stem characteristics in the maize–soybean strip intercropping system. Frontiers in Plant Science, 13, 848893.

Chi B J, Liu J, Dai J L, Li Z H, Zhang D M, Xu S Z, Nie J J, Wan S M, Li C D, Dong H Z. 2023. Alternate intercropping of cotton and peanut increases productivity by increasing canopy photosynthesis and nutrient uptake under the influence of rhizobacteria. Field Crops Research, 302, 109059.

Czaban W, Han E, Lund O S, Stokholm M S, Jensen S M, Thorup-Kristensen K. 2023. The enhancing effect of intercropping sugar beet with chicory on the deep root growth and nutrient uptake. Agriculture, Ecosystems & Environment, 347, 108360.

Falster D S, Westoby M. 2003a. Leaf size and angle vary widely across species: What consequences for light interception?. New Phytologist, 158, 509-525.

Falster D S, Westoby M. 2003b. Plant height and evolutionary games. Trends in Ecology & Evolution, 18, 337-343.

Gu Y, Zheng H Y, Li S, Wang W T, Guan Z Y, Li J Z, Mei N, Hu W H. 2024. Effects of narrow-wide row planting patterns on canopy photosynthetic characteristics, bending resistance and yield of soybean in maize‒soybean intercropping systems. Scientific Reports, 14, 9361.

He S Y, Li X N, Chen M G, Xu X Y, Zhang W J, Chi H L, Shao P X, Tang F D, Gong T, Guo M, Xu M, Yang W Y, Liu W G. 2024. Excellent canopy structure in soybeans can improve their photosynthetic performance and increase yield. Agriculture, 14, 1783.

Hu J, Zhao X Y, Gu L M, Liu P, Zhao B, Zhang J W, Ren B Z. 2023. The effects of high temperature, drought, and their combined stresses on the photosynthesis and senescence of summer maize. Agricultural Water Management, 289, 108525.

Jiang P, Wang Y Z, Zhang Y P, Fei J C, Rong X M, Peng J W, Yin L C, Luo G W. 2024. Intercropping enhances maize growth and nutrient uptake by driving the link between rhizosphere metabolites and microbiomes. New Phytologist, 243, 1506-1521.

Jing B, Shi W J, Wang H, Lin F M. 2023. 15N labeling technology reveals enhancement of nitrogen uptake and transfer by root interaction in cotton/soybean intercropping. Journal of the Science of Food and Agriculture, 103, 6307-6316.

Jolliffe P A, Wanjau F M. 1999. Competition and productivity in crop mixtures: Some properties of productive intercrops. The Journal of Agricultural Science, 132, 425-435.

Kandel B P. 2020. Spad value varies with age and leaf of maize plant and its relationship with grain yield. BMC Research Notes, 13, 475.

Khan A, Wang L S, Ali S, Tung S A A, Hafeez A, Yang G Z. 2017. Optimal planting density and sowing date can improve cotton yield by maintaining reproductive organ biomass and enhancing potassium uptake. Field Crops Research, 214, 164-174.

Khan M A, Wahid A, Ahmad M, Tahir M T, Ahmed M, Ahmad S, Hasanuzzaman M. 2020. World cotton production and consumption: An overview. In: Cotton Production and Uses: Agronomy, Crop Protection, and Postharvest Technologies. Springer, Singapore. pp. 1-7.

Kong W S, Wei M, Khan N, Liang J, Han D Q, Zhang H J. 2024. Assessing sustainable future of import-independent domestic soybean production in China: Policy implications and projections for 2030. Frontiers in Sustainable Food Systems, 8, 1387609.

Kou H T, Liao Z Q, Liu Y Y, Lai Z L, Li Z J, Fan J L. 2025a. Row configuration affects plant growth and grain yield of drip-irrigated maize-soybean strip intercropping system by regulating photosynthetic capacity and grain filling process. Journal of Agriculture and Food Research, 21, 101920.

Kou H T, Liao Z Q, Liu Y Y, Zhang H, Lai Z L, Yu J, Li Z J, Fan J L. 2025b. Light distribution, interception and use efficiency of drip-fertigated maize-soybean strip intercropping systems under various row configuration conditions. Plant, Cell & Environment.

Li C J, Stomph T J, Makowski D, Li H G, Zhang C C, Zhang F S, Van Der Werf W. 2023. The productive performance of intercropping. Proceedings of the National Academy of Sciences of the United States of America, 120, e2201886120.

Li J, Xie R Z, Wang K R, Hou P, Ming B, Zhang G Q, Liu G Z, Wu M, Yang Z S, Li S K. 2018. Response of canopy structure, light interception and grain yield to plant density in maize. The Journal of Agricultural Science, 156, 785-794.

Li Y H, Shi D Y, Li G H, Zhao B, Zhang J W, Liu P, Ren B Z, Dong S T. 2019. Maize/peanut intercropping increases photosynthetic characteristics, ¹³C-photosynthate distribution, and grain yield of summer maize. Journal of Integrative Agriculture, 18, 2219-2229.

Li Y J, Ma L S, Wu P T, Zhao X N, Chen X L, Gao X D. 2020. Yield, yield attributes and photosynthetic physiological characteristics of dryland wheat (Triticum aestivum L.)/maize (Zea mays L.) strip intercropping. Field Crops Research, 248, 107656.

Liang J P, He Z J, Shi W J. 2020. Cotton/mung bean intercropping improves crop productivity, water use efficiency, nitrogen uptake, and economic benefits in the arid area of Northwest China. Agricultural Water Management, 240, 106277.

Liu X, Rahman T, Song C, Yang F, Su B Y, Cui L, Bu W Z, Yang W Y. 2018. Relationships among light distribution, radiation use efficiency and land equivalent ratio in maize-soybean strip intercropping. Field Crops Research, 224, 91-101.

Lu J T, Dong Q Q, Lan G H, He Z C, Zhou D Y, Zhang H, Wang X G, Liu X B, Jiang C J, Zhang Z, Wan S B, Zhao X H, Yu H Q. 2023. Row ratio increasing improved light distribution, photosynthetic characteristics, and yield of peanut in the maize and peanut strip intercropping system. Frontiers in Plant Science, 14, 1135580.

Lv Q Q, Chi B J, He N, Zhang D M, Dai J L, Zhang Y J, Dong H Z. 2023. Cotton-based rotation, intercropping, and alternate intercropping increase yields by improving root–shoot relations. Agronomy, 13, 413.

Lv Q Q, Dai J L, Ding K D, He N, Li Z H, Zhang D M, Xu S Z, Li C D, Chi B J, Zhang Y J, Dong H Z. 2024. Managing interspecific competition to enhance productivity through selection of soybean varieties and sowing dates in a cotton-soybean intercropping system. Field Crops Research, 316, 109513.

Lv Q Q, He N, Chi B J, Xu S Z, Li Z H, Wang L, Zhang Y J, Zhang D M, Cui Z P, Dai J L, Nie J J, Zhang Y J, Gan Y T, Dong H Z. 2025. Spatiotemporal diversification enables sustainable cotton-soybean production with enhanced yield and reduced emissions. Journal of Cleaner Production, 527, 146709.

Mai W X, Xue X R, Gu F, Tian C Y. 2018. Simultaneously maximizing root/mycorrhizal growth and phosphorus uptake by cotton plants by optimizing water and phosphorus management. BMC Plant Biology, 18, 334.

Mead R, Willey R. 1980. The concept of a ‘land equivalent ratio’and advantages in yields from intercropping. Experimental Agriculture, 16, 217-228.

Nasar J, Shao Z, Arshad A, Jones F G, Liu S, Li C, Khan M Z, Khan T, Banda J S K, Zhou X, Gao Q. 2020. The effect of maize–alfalfa intercropping on the physiological characteristics, nitrogen uptake and yield of maize. Plant Biology, 22, 1140-1149.

Pagano M C, Miransari M. 2016. The importance of soybean production worldwide. In: Abiotic and Biotic Stresses in Soybean Production. Academic Press. America. pp. 1-26.

Parry M A J, Reynolds M, Salvucci M E, Raines C, Andralojc P J, Zhu X G, Price G D, Condon A G, Furbank R T. 2011. Raising yield potential of wheat. II. Increasing photosynthetic capacity and efficiency. Journal of Experimental Botany, 62, 453-467.

Qiang B B, Fan Z, Tang N, Asad M S, Timbang B C, Ren X L, Chen X L. 2024. Improving the productivity of intercropping through above and below ground separation: A case study on photosynthetic characteristics and root distribution. Industrial Crops and Products, 222, 119506.

Qiao M J, Sun R B, Wang Z X, Dumack K, Xie X G, Dai C C, Wang E T, Zhou J Z, Sun B, Peng X H, Bonkowski M, Chen Y. 2024. Legume rhizodeposition promotes nitrogen fixation by soil microbiota under crop diversification. Nature Communications, 15, 2924.

Qiu Y R, Li X, Tang Y R, Xiong S W, Han Y C, Wang Z B, Feng L, Wang G P, Yang B F, Lei Y P, Du W L, Zhi X Y, Xin M H, Jiao Y H, Zhang S J, Li Y B, Li X F. 2023. Directly linking plant N, P and K nutrition to biomass production in cotton-based intercropping systems. European Journal of Agronomy, 151, 126960.

Raza M A, Din A M U, Yasin H S, Gul H, Saeed A, Mehmood A, Rehman S U, Iqbal Z, Iqbal R, Kubaisi N A, Elshikh M S, Shah G A, Wang Z Q, Rahman M H U, Khalid M H B, Haider I, Ma Z M. 2025. Yield gains and resource use advantages driven by legume choice and row ratio in cotton/legume intercropping under arid-irrigated conditions. Field Crops Research, 324, 109789.

Raza M A, Wang Z Q, Yasin H S, Gul H, Qin R J, Rehman S U, Mahmood A, Iqbal Z, Ahmed Z, Luo S L, Chen J, Liang X, Gitari H, Khalid M H B, Feng Y, Ma Z M. 2023. Effect of crop combination on yield performance, nutrient uptake, and land use advantage of cereal/legume intercropping systems. Field Crops Research, 304, 109144.

Raza M A, Yasin H S, Gul H, Qin R J, Din A M U, Khalid M H B, Hussain S, Gitari H, Saeed A, Wang J, Chiyaneh E R, Sabagh A E, Manzoor A, Fatima A, Ahmad S, Yang F, Skalicky M, Yang W Y. 2022. Maize/soybean strip intercropping produces higher crop yields and saves water under semi-arid conditions. Frontiers in Plant Science, 13, 1006720.

Ren Y Y, Zhang L, Yan M F, Zhang Y J, Chen Y L, Palta J A, Zhang S Q. 2021. Effect of sowing proportion on above-and below-ground competition in maize–soybean intercrops. Scientific Reports, 11, 15760.

Salama H S A, Abdel-Moneim M H. 2021. Maximizing land use efficiency and productivity of soybean and fodder maize intercrops through manipulating sowing schedule and maize harvest regime. Agronomy, 11, 863.

Schwerdtner U, Spohn M. 2022. Plant species interactions in the rhizosphere increase maize N and P acquisition and maize yields in intercropping. Journal of Soil Science and Plant Nutrition, 22, 3868-3884.

Smith E N, van Aalst M, Tosens T, Niinemets Ü, Stich B, Morosinotto T, Alboresi A, Erb T J, Gómez-Coronado P A, Tolleter D, Finazzi G, Curien G, Heinemann M, Ebenhöh O, Hibberd J M, Schlüter U, Sun T, Weber A P M. 2023. Improving photosynthetic efficiency toward food security: Strategies, advances, and perspectives. Molecular Plant, 16, 1547-1563.

Tang Z J, Wang X, Xiang Y Z, Liang J P, Guo J J, Li W Y, Lu J S, Du R Q, Li Z J, Zhang F C. 2024. Application of hyperspectral technology for leaf function monitoring and nitrogen nutrient diagnosis in soybean (Glycine max L.) production systems on the Loess Plateau of China. European Journal of Agronomy, 154, 127098.

Tian X L, Wang C B, Bao X G, Wang P, Li X F, Yang S C, Ding G C, Christie P, Li L. 2019. Crop diversity facilitates soil aggregation in relation to soil microbial community composition driven by intercropping. Plant and Soil, 436, 173-192.

Wang C, Zhou L B, Zhang G B, Gao J, Peng F L, Zhang C L, Xu Y, Zhang L Y, Shao M B. 2021. Responses of photosynthetic characteristics and dry matter formation in waxy sorghum to row ratio configurations in waxy sorghum-soybean intercropping systems. Field Crops Research, 263, 108077.

Wang G F, Wang D P, Zhou X Y, Shah S, Wang L C, Ahmed M, Sayyed R Z, Fahad S. 2022. Effects of cotton–peanut intercropping patterns on cotton yield formation and economic benefits. Frontiers in Sustainable Food Systems, 6, 900230.

Wang J B, Chen G D, Wang P J, Cui Z J, Wan S M, Zhai Y L, Li T T, Zhao Y R. 2024. Optimizing cotton row configuration in jujube–cotton intercropping systems improves their productivity, net effects, and sustainability. Agronomy, 14, 1216.

Wang L, Zhou T, Cheng B, Du Y L, Qin S S, Gao Y, Xu M, Lu J J, Liu T, Li S X, Liu W G, Yang W Y. 2020. Variable light condition improves root distribution shallowness and P uptake of soybean in maize/soybean relay strip intercropping system. Plants, 9, 1204.

Wang R N, Sun Z X, Bai W, Wang E L, Wang Q, Zhang D S, Zhang Y, Yang N, Liu Y, Nie J Y, Chen Y F, Duan L S, Zhang L Z. 2021. Canopy heterogeneity with border-row proportion affects light interception and use efficiency in maize/peanut strip intercropping. Field Crops Research, 271, 108239. 

Wang Y Z, Zhang Y P, Yang Z Y, Fei J C, Zhou X, Rong X M, Peng J W, Luo G W. 2024. Intercropping improves maize yield and nitrogen uptake by regulating nitrogen transformation and functional microbial abundance in rhizosphere soil. Journal of Environmental Management, 358, 120886.

Wang Z K, Zhao X N, Wu P T, He J Q, Chen X L, Gao Y, Cao X C. 2015. Radiation interception and utilization by wheat/maize strip intercropping systems. Agricultural and Forest Meteorology, 204, 58-66.

Wu G, Yu F F, Yuan M M, Wang J B, Liu C, He W Z, Ge Z H, Sun Y X, Liu Y. 2022. Responses of rhizosphere bacterial and fungal communities to the long-term continuous monoculture of water oat. Microorganisms, 10, 2174.

Wu Y S, Chen M, Huang S R, Li Y, Li M, He D, Hu P C, Duan T, Gong W Z, Yan Y H, Kwame T J, Raza M A, Yang W Y. 2025. Combining modelling and experiment to quantify light interception and inter row variability on intercropped soybean in strip intercropping. European Journal of Agronomy, 164, 127508.

Xi H, Shen J L, Qu Z, Yang D Y, Liu S M, Nie X H, Zhu L F. 2019. Effects of long-term cotton continuous cropping on soil microbiome. Scientific Reports, 9, 18297.

Xiao S S, Ming L W, Zhang Y F, Wang Z Y, Li F M, Wang T H, Zhang C Y, Yang K J, Yu S, Li M K, Yu S Q, Hou J J, An J Y, Guo M J, Tian X J, Liu J H. 2025. Effects of intercropping long- and short-season varieties on the photosynthetic characteristics and yield formation of maize in high-latitude cold regions. Agronomy, 15, 2505.

Xie W, Zhang K, Wang X Y, Zou X X, Zhang X J, Yu X N, Wang Y F, Si T. 2022. Peanut and cotton intercropping increases productivity and economic returns through regulating plant nutrient accumulation and soil microbial communities. BMC Plant Biology, 22, 121.

Yin W, Chai Q, Zhao C, Yu A Z, Fan Z L, Hu F L, Fan H, Guo Y, Coulter J A. 2020. Water utilization in intercropping: A review. Agricultural Water Management, 241, 106335.

Yu Y, Stomph T J, Makowski D, Van der Werf W. 2015. Temporal niche differentiation increases the land equivalent ratio of annual intercrops: A meta-analysis. Field Crops Research, 184, 133-144.

Zhai M H, Wei X W, Pan Z L, Xu Q Q, Qin D L, Li J H, Zhang J, Wang L Z, Wang K F, Duan X Y, Zhang Y P, Zhao W Q, Li A, Zhang Z G, Wang Z B. 2024. Optimizing plant density and canopy structure to improve light use efficiency and cotton productivity: Two years of field evidence from two locations. Industrial Crops and Products, 222, 119946.

Zhang D S, Zhang L Z, Liu J G, Han S, Wang Q, Evers J, Liu J, Van der Werf W, Li L. 2014. Plant density affects light interception and yield in cotton grown as companion crop in young jujube plantations. Field Crops Research, 169, 132-139.

Zhang L, Van der Werf W, Zhang S, Li B, Spiertz J H J. 2007. Growth, yield and quality of wheat and cotton in relay strip intercropping systems. Field Crops Research, 103, 178-188.

Zhang S T, Wang J B, Cui Z J, Li T T, Dong Z L, Qiao H, Li L, Wan S M, Li X F, Zhang W, Hu Q, Chen G D. 2025. Jujube–cotton intercropping enhances yield and economic benefits via photosynthetic regulation in oasis agroecosystems of southern Xinjiang. Agronomy, 15, 1676.

Zhang Y T, Liu J, Zhang J Z, Liu H B, Liu S, Zhai L M, Wang H Y, Lei Q L, Ren T Z, Yin C B. 2015. Row ratios of intercropping maize and soybean can affect agronomic efficiency of the system and subsequent wheat. PLoS ONE, 10, e0129245.