Bardgett R, Van der Putten W. 2014. Belowground biodiversity and ecosystem functioning. Nature, 515, 505–511.

Chen G P, Liu M, Zhao X Y, Bawa G, Liang B, Feng L, Pu T, Yong T W, Liu W G, Liu J, Du J B, Yang F, Wu Y S, Liu C Y, Wang X C, Yang W Y. 2023. Improved photosynthetic performance under unilateral weak light conditions in a wide–narrow-row intercropping system is associated with altered sugar transport. Journal of Experimental Botany, 75, 258-273.

Chen P, Xian G J, Pu T, Gong W Z, Li J Z, Yang F, Wang X C, Yong T W, Li Y Z, Yan Y H, Liu J, Yang W Y, Wu Y S. 2025. Soybean-maize intercropping combined with water management enhances leaf traits to obtain yield benefits in semi-arid regions. Field Crops Research, 327, 109869.

Chen X Y, Ren H, Zhang J W, Zhao B, Ren B Z, Wan Y S, Liu P. 2024. Deep phosphorus fertilizer placement increases maize productivity by improving root-shoot coordination and photosynthetic performance. Soil and Tillage Research, 235, 105915.

David R, Mark A S, Omar S, Ralph B. 2024. Structure, function, and assembly of PSI in thylakoid membranes of vascular plants. Plant Cell, 36, 4080-4108.

Du J B, Han T F, Gai J Y, Yong T W, Sun X, Wang X C, Yang F, Liu J, Shu K, Liu W G, Yang W Y. 2018. Maize-soybean strip intercropping: Achieved a balance between high productivity and sustainability. Journal of Integrative Agriculture, 17, 747-754.

Feng L, Chen G P, Pu T, Liang B, Liu X, Wu Y S, Yong T W, Yang F, Wang X C, Yang W Y. 2024. Analyzing the stable yield mechanism of maize-soybean intercropping based on the perspective of assimilation product distribution, leaf photosynthetic physiology and leaf anatomical structure. Plant and Soil, 504, 367-384.

Gu Y R, Ibitoye R G, He J J, Gao F, Zhou X B. 2025. Photosynthetic performance and yield of intercropped maize and soybean are directly opposite under different intercropping ratios and maize planting densities interactions. Journal of the Science of Food and Agriculture, 105, 7440-7452.

Guan C L, Zhang D, Chu C C. 2025. Interplay of light and nitrogen for plant growth and development. The Crop Journal, 13, 641-655.

Huang Y R, Wang M Q, Wei L L, Zhao C J, Yang L, Zhou X B. 2025. Optimizing photosynthesis, resource use efficiency, and yield in maize-soybean intercropping through row configurations in Southwest China. Plant Biology, 27, 1202-1213.

Li Y L, Chen P, Fu Z D, Luo K, Lin P, Gao C, Liu S S, Pu T, Yong T W, Yang W Y. 2023. Maize-soybean relay cropping increases soybean yield synergistically by extending the post-anthesis leaf stay-green period and accelerating grain filling. The Crop Journal, 11, 1921-1930.

Liang B, Ma Y W, Shi K, Chen G P, Chen H, Hu Y, Chen P, Pu T, Wu Y S, Sun X, Yong T W, Liu W G, Liu J, Du J B, Yang F, Wang X C, Yang W Y. 2023. Appropriate bandwidth achieves a high yield by reducing maize intraspecific competition in additive maize–soybean strip intercropping. European Journal of Agronomy, 142, 126658.

Liao Z Q, Zhang C, Zhang Y, Yu S L, Yan S C, Zhang S H, Li Z J, Fan J L. 2024. Nitrogen application and soil mulching improve grain yield of rainfed maize by optimizing source-sink relationship and grain filling process on the Loess Plateau of China. European Journal of Agronomy, 153, 127060.

Liu C H, Pang S Y, Li X F, Liu P, Zhou Y, Lin X, Gu S B, Wang D. 2025. Layered nitrogen fertilization regulates root morphology to promote synergistic nitrogen and phosphorus absorption in maize (Zea mays L.). Field Crops Research, 322, 109737.

Liu G Z, Yang Y S, Guo X X, Liu W M, Xie R Z, Ming B, Xue J, Wang K R, Li S K, Hou P. 2022. Coordinating maize source and sink relationship to achieve yield potential of 22.5 Mg ha^-1. Field Crops Research, 283, 108544.

Liu J, Yang W Y. 2024. Soybean maize strip intercropping: A solution for maintaining food security in China. Journal of Integrative Agriculture, 23, 2503-2506.

Mason T G, Maskell E J. 1928. Studies on the transport of carbohydrates in the cotton plant: II. the factors determining the rate and the direction of movement of sugars. Annals of Botany, 42, 571–636.

Nasar J, Zhao C J, Khan R, Gul H, Gitari H, Shao Z, Abbas G, Haider I, Iqbal Z, Ahmed W, Rehman R, Liang Q P, Zhou X B, Yang J. 2023. Maize-soybean intercropping at optimal N fertilization increases the N uptake, N yield and N use efficiency of maize crop by regulating the N assimilatory enzymes. Frontiers in Plant Science, 13, 1077948.

Ren H, Liu Z, Wang X B, Zhou W B, Zhou B Y, Zhao M, Li C F. 2025. Long-term excessive nitrogen application decreases spring maize nitrogen use efficiency via suppressing root physiological characteristics. Journal of Integrative Agriculture, 24, 4195-4210.

Rodriguez C, Carlsson G, Englund J, Adam F, Pelzer E, Jeuffroy M, Makowski D, Jensen E S. 2020. Grain legume-cereal intercropping enhances the use of soil-derived and biologically fixed nitrogen in temperate agroecosystems. A meta-analysis. European Journal of Agronomy, 118, 126077.

Tilman D, Cassman K G, Matson P A, Naylor R, Polasky S. 2002. Agricultural sustainability and intensive production practices. Nature, 418, 671–677.

Wu P, Chen G Z, Liu F, Cai T, Zhang P, Jia Z K. 2021. How does deep-band fertilizer placement reduce N₂O emissions and increase maize yields? Agriculture, Ecosystems & Environment, 322, 107672.

Wu P, Liu F, Wang J Y, Liu Y H, Gao Y, Zhang X Q, Chen G Z, Huang F Y, Ahmad S, Zhang P, Cai T, Jia Z K. 2022. Suitable fertilization depth can improve the water productivity and maize yield by regulating development of the root system. Agricultural Water Management, 271, 1077784.

Xiao T, Din A, Cui K S, Raza M A, Ali M F, Xiao J H. 2023. Inter-specific root interactions and water use efficiency of maize/soybean relay strip intercropping. Field Crops Research, 291, 108793.

Yan Y Y, Duan F Y, Li X, Zhao R L, Hou P, Zhao M, Li S K, Wang Y H, Dai T B, Zhou W B. 2024. Photosynthetic capacity and assimilate transport of the lower canopy influence maize yield under high planting density. Plant Physiology, 195, 2652-2667.

Yang H, Su Y H, Wang L, Whalen J K, Pu T, Wang X C, Yang F, Yong T W, Liu J, Yan Y H, Yang W Y, Wu Y S. 2025. Strip intercropped maize with more light interception during post-silking promotes photosynthesized carbon sequestration in the soil. Agriculture Ecosystems & Environment, 378, 109301.

Yu R P, Yang H, Xing Y, Zhang W P, Lambers H, Li L. 2022. Belowground processes and sustainability in agroecosystems with intercropping. Plant and Soil, 476, 263-288.

Zeng H Q, Chen H Y, Zhang M X, Ding M, Xu F Y, Yan F, Toshinori K, Zhu Y Y. 2024. Plasma membrane H⁺-ATPases in mineral nutrition and crop improvement. Trends in Plant Science, 29, 978-994.

Zhai Z M, Shi W J, Liu L, Jing B. 2024. Effects of poly-γ-glutamic acid on soil pore structure, maize grain filling, yield and water use efficiency under mulched drip irrigation. Field Crops Research, 317, 109528.

Zhang D S, Sun Z X, Feng L S, Bai W, Yang N, Zhang Z, Du G J, Feng C, Cai Q, Wang Q, Zhang Y, Wang R N, Adnan A, Hao X Y, Sun M, Gao Z Q, Zhang L Z. 2020. Maize plant density affects yield, growth and source-sink relationship of crops in maize/peanut intercropping. Field Crops Research, 257, 107926.

Zhang Y, Sun Z X, Su Z C, Du G J, Bai W, Wang Q, Wang R N, Nie J Y, Sun T R, Feng C, Zhang Z, Yang N, Zhang X, Evers J, Wopke V, Zhang L Z. 2022. Root plasticity and interspecific complementarity improve yields and water use efficiency of maize/soybean intercropping in a water-limited condition. Field Crops Research, 282, 108523.

Zhao X Y, Hu Y, Liang B, Chen G P, Feng L, Pu T, Sun X, Yong T W, Liu W G, Liu J, Du J B, Yang F, Wang X C, Yang W Y. 2023. Coordination of density and nitrogen fertilization improves stalk lodging resistance of strip intercropped maize with soybeans by affecting stalk quality traits. Agriculture, 13, 13051009.

Zhu Y P, Wang S, Li Y H, Wei D, Luo N, Wang P, Meng Q F. 2023. Enhancing maize yield through understanding the novel shoot-root interaction in morphology among hybrids with dense planting. Field Crops Research, 302, 109107.