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Journal of Integrative Agriculture  2026, Vol. 25 Issue (7): 3017-3030    DOI: 10.1016/j.jia.2025.09.011
Agro-ecosystem & Environment Advanced Online Publication | Current Issue | Archive | Adv Search |
Optimizing maize yield and kernel quality via leguminous green manure intercropping with deficit irrigation in arid agroecosystem

Diaoliang Zhang1, Yunyou Nan2, Zhilong Fan1, 2#, Qiang Chai1, 2#, Gary Y. Gan1, 3#, Wen Yin1, 2, Falong Hu2

1 State Key Laboratory of Arid Land Crop Science, Gansu Agricultural University, Lanzhou 730070, China

2 College of Agronomy, Gansu Agricultural University, Lanzhou 730070, China

3 College of Life and Environmental Sciences, Wenzhou University, Wenzhou 325035, China

 Highlights 
In leguminous green manure intercropping system, 15% deficit irrigation can achieve maize yield equivalent to conventional irrigation.
It also enhanced kernel quality and sustained high levels of protein and nutrients under deficit irrigation.
These improvements were attributed to an increased net photosynthetic rate and a greater soil nitrogen supply.
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摘要  

间作豆科绿肥是一种通过提升土壤肥力和资源利用效率来增强农业生态系统韧性的可持续策略。然而,在干旱条件下,间作豆科绿肥与调亏灌溉协同维持玉米产量稳定性并改善籽粒品质的机制仍不明晰。本研究开展了一项为期三年(2021–2023年)的田间裂区试验,主区设3种绿肥还田方式:绿肥全量翻压还田M||V-P)、绿肥还田M||V-R)以及无绿肥对照(单作玉米,SM);副区设3种灌溉制度:常规灌溉(I3400 mm)、15%亏缺灌溉(I2340 mm)和30%亏缺灌溉(I1280 mm)。试验测定了玉米籽粒产量、籽粒品质(蛋白质、脂肪、淀粉及必需氨基酸含量)、玉米净光合速率(Pn)以及土壤硝态氮与铵态氮含量结果表明,与单作玉米(SM)相比,M||V-P M||V-R 处理均提高了玉米籽粒产量,其中 M||V-P 的产量比 M||V-R 高出 5.7%。值得注意的是,M||V-P I215%亏缺灌溉)条件下的产量与 M||V-P I3(常规灌溉)条件下相当,且较 SMI3(单作玉米常规灌溉)增产 18.3%。与单作玉米相比,M||V-P M||V-R 还显著提升了籽粒品质,表现为蛋白质、脂肪、淀粉及必需氨基酸含量增加。随着灌溉量减少,籽粒蛋白质含量有所增加,而脂肪和淀粉含量降低。特别地,M||V-P I2 条件下的籽粒蛋白质含量与 I130%亏缺灌溉)处理无显著差异,同时脂肪、淀粉及必需氨基酸含量与 I3(常规灌溉)处理相近。与 SMI3 相比,M||V-P I2 条件下所有籽粒品质指标均显著增加这些改善主要得益于间作豆科绿肥在减少 15% 灌溉用水的同时,使玉米净光合速率(Pn)提高了 14.3%,并将土壤硝态氮与铵态氮含量分别提升了 12.5% 5.2%。本研究表明,间作豆科绿肥结合调亏灌溉可作为水资源受限地区玉米可持续生产的一种推广模式。



Abstract  

Intercropping with leguminous green manure represents a sustainable approach to enhance agroecosystem resilience through improved soil fertility and resource-use efficiency.  However, the synergistic mechanisms between leguminous green manure intercropping and regulated deficit irrigation in maintaining maize yield stability and enhancing kernel profiles under arid conditions remain inadequately understood.  A three-year (2021–2023) split-plot field experiment incorporated main plots consisting of three green manure incorporation practices: full green manure incorporation (M||V-P), green manure stubble retention (M||V-R), and maize without green manure (maize sole cropping, SM); while split plots comprised three irrigation regimes: conventional (I3; 400 mm), 15% deficit (I2; 340 mm), and 30% deficit (I1; 280 mm).  The study examined maize grain yield, kernel quality (protein, fat, starch, and essential amino acid content), net photosynthetic rate (Pn) of maize, and soil nitrate-ammonium nitrogen content.  M||V-P and M||V-R increased maize grain yield compared to SM, with M||V-P producing 5.7% higher yields than M||V-R.  Notably, M||V-PI2 achieved comparable yield to M||V-PI3 while reducing irrigation by 15%, demonstrating an 18.3% yield increase over SMI3.  M||V-P and M||V-R enhanced kernel quality compared to SM, exhibiting higher protein, fat, starch, and essential amino acid content.  Decreased irrigation led to increased kernel protein content but reduced fat and starch contents.  The kernel protein content under M||V-PI2 showed no significant difference from M||V-PI1, while maintaining fat, starch, and essential amino acid content similar to M||V-PI3.  M||V-PI2 improved all kernel quality parameters relative to SMI3.  These enhancements primarily resulted from maize intercropped with leguminous green manure in combination with 15% deficit irrigation, which increased maize Pn by 14.3%, and elevated soil nitrate-ammonium nitrogen by 12.5 and 5.2%, respectively.  These findings demonstrate a scalable approach for sustainable maize production though the integration of leguminous green manure intercropping in water-limited regions.

Keywords:  intercropping with leguminous green manure       yield–water tradeoff       kernel quality       maize  
Received: 21 April 2025   Accepted: 08 August 2025 Online: 09 September 2025  
Fund: 

We appreciate the support from the National Natural Science Foundation of China (32160765) and the Ear-marked Fund for China Agriculture Research System (CARS-22-G-12).

About author:  Diaoliang Zhang, Mobile: +86-18809405878, E-mail: zhangdl0320@163.com; #Correspondence Zhilong Fan, Mobile: +86-18919113084, E-mail: fanzl@gsau.edu.cn; Qiang Chai, Tel: +86-931-7631145; E-mail: chaiq@gsau.edu.cn; Gary Y. Gan, E-mail: gary.gan@wzu.edu.cn

Cite this article: 

Diaoliang Zhang, Yunyou Nan, Zhilong Fan, Qiang Chai, Gary Y. Gan, Wen Yin, Falong Hu. 2026. Optimizing maize yield and kernel quality via leguminous green manure intercropping with deficit irrigation in arid agroecosystem. Journal of Integrative Agriculture, 25(7): 3017-3030.

Alqudah A M, Samarah N H, Mullen R E. 2011. Drought stress effect on crop pollination, seed set, yield and quality. Sustainable Agriculture Review6, 193–213.

Anugroho F, Kitou M, Nagumo F, Kinjo K, Jayasinghe G Y. 2010. Potential growth of hairy vetch as a winter leguminous cover crops in subtropical soil conditions. Soil Science and Plant Nutrition56, 254–262.

Chen Y N, Fang G H, Li Z, Zhang X Q, Gao L, Elbeltagi A, Shaer H E, Duan W L, Wassif O M A, Li Y P, Luo P P, Selmi A, Yu R D, Yang J H, Hu Y N, Liu C X, Long Y X, Malik I, Fu A H, Wistuba M, et al. 2024. The crisis in oases: Research on ecological security and sustainable development in arid regions. Annual Review of Environment and Resources49, 1–20.

Cheng G D, Kong F X, Gan Y T, Zhang R Z, Feng F X, Yu A Z, Zhao C, Wan S M, Chai Q. 2018. Enhancing the systems productivity and water use efficiency through coordinated soil water sharing and compensation in strip-intercropping. Scientific Reports8, 10494.

D’Amzto R, Feudis D M, Guiducci M, Businelli D. 2019. Zea mays L. grain: Increase in nutraceutical and antioxidant properties due to Se fortification in low and high-water regimes. Journal of Agricultural and Food Chemistry67, 7050–7059.

Daryanto S, Fu B J, Wang L X, Jacinthe P A, Zhao W W. 2018. Quantitative synthesis on the ecosystem services of cover crops. Earth-Science Reviews185, 357–373.

De Santis M A, Soccio M, Laus M N, Flagella Z. 2021. Influence of drought and salt stress on durum wheat grain quality and composition: A review. Plants10, 2599.

Fan Z L, Chai Q, Yu A Z, Zhao C, Yin W, Hu F L, Chen G P, Cao W D, Coulter J A. 2020. Water and radiation use in maize–pea intercropping is enhanced with increased plant density. Agronomy Journal112, 257–273.

Feng W J, Xue W W, Zhao Z Q, Shi Z K, Wang W J, Bai Y, Wang H X, Qiu P, Xue J F, Chen B G. 2024. Nitrogen fertilizer application rate affects the dynamic metabolism of nitrogen and carbohydrates in kernels of waxy maize. Frontiers in Plant Science15, 1416397.

Franco J G, King S R, Volder A. 2018. Component crop physiology and water use efficiency in response to intercropping. European Journal of Agronomy93, 27–39.

Gao H X, Zhang C C, Van Der Werf W, Ning P, Zhang Z, Wan S B, Zhang F S. 2022. Intercropping modulates the accumulation and translocation of dry matter and nitrogen in maize and peanut. Field Crops Research284, 108561.

Gong C, Tan Q Y, Xu M X, Liu G B. 2020. Mixed-species plantations can alleviate water stress on the loess plateau. Forest Ecology and Management458, 117767.

Guo J, Qu L L, Wang L F, Lu W P, Lu D L. 2023. Effects of post-silking drought stress degree on grain yield and quality of waxy maize. Journal of the Science of Food and Agriculture103, 1530–1540.

Hou S, Sun X R, Chen G H, Siddique K H, Chen Z L, Liu F, Ping S Y, Lai H T, Guo H H, An Y J, Lin Z L, Zhang Z X, Sun L Z, Yang P Z. 2024. Biochar addition mitigates asymmetric competition of water and increases yield advantages of maize–alfalfa strip intercropping systems in a semiarid region on the Loess Plateau. Field Crops Research319, 109645.

Hu F L, Tan Y, Yu A Z, Zhao C, Coulter J A, Fan Z L, Yin W, Fan H, Chai Q. 2018. Low N fertilizer application and intercropping increases N concentration in pea (Pisum sativum L.) grains. Frontiers in Plant Science9, 1763.

Hu Z H, Zhao Q, Zhang X J, Ning X G, Liang H, Cao W D. 2023. Winter green manure decreases subsoil nitrate accumulation and increases N use efficiencies of maize production in North China Plain. Plants12, 311.

Iderawumi A M, Kamal T O. 2022. Green manure for agricultural sustainability and improvement of soil fertility. Farming and Management7, 1–8.

Jing B, Shi W J, Chen T, Zhai Z M, Song J W. 2025. Optimizing root distribution and water use efficiency in maize/soybean intercropping under different irrigation levels: The role of underground interactions. Soil and Tillage Research249, 106490.

Kashiwagi T. 2021. Effects of rice grain protein QTL, TGP12, on grain composition, yield components, and eating quality with different nitrogen applications. Field Crops Research263, 108051.

Li C J, Stomph T J, Makowski D, Li H G, Zhang C C, Zhang F S, Van Der Werf W. 2023. The productive performance of intercropping. Proceedings of the National Academy of Sciences of the United States of America120, e2201886120.

Li H R, Li X L, Mei X R, Nangia V, Guo R, Hao W P, Wang J D. 2023. An alternative water-fertilizer-saving management practice for wheat–maize cropping system in the North China Plain: Based on a 4-year field study. Agricultural Water Management276, 108053.

Li H T, Wang G C, Hu F L, Fan Z L, Yin W, Cao W D, Chai Q, Yao T. 2025. Additive intercropping green manure enhances maize water productivity through water competition and compensation under reduced nitrogen fertilizer application. Agricultural Water Management311, 109388.

Li J Y, Wen X J, Yang J Z, Yang W Y, Xin Y F, Zhang L, Liu H P, He Y L, Yan Y H. 2022. Effects of maize varieties on biomass yield and silage quality of maize–soybean intercropping in the Qinghai-Tibet Plateau. Fermentation8, 542.

Li J Z, Li H S, Cai Y, Li B, He C, Li Z, Zheng Z P. 2023. Effects of planting and turning green manure in winter under nitrogen fertilizer reduction on corn grain quality. Journal of Sichuan Agricultural University41, 393–398, 415. (in Chinese)

Li P, Jia L, Chen Q Q, Zhang H J, Deng J J, Lu J Y, Xu L, Li H X, Hu F, Jiao J G. 2024. Adaptive evaluation for agricultural sustainability of different fertilizer management options for a green manure–maize rotation system: Impacts on crop yield, soil biochemical properties and organic carbon fractions. Science of the Total Environment908, 168170.

Li X F, Wang Z G, Wu J P, Liu X R, Tian X L, Wang Y, Li J P, Wang Y, Xia H Y, Mei P P, Wang X F, Yu R P, Zhang W P, Li L, Bao X G, Sun J H, Yang S C, Wang C B, Zhao J H, Che Z X, et al. 2021. Long-term increased grain yield and soil fertility from intercropping. Nature Sustainability11, 4.

Liu H J, Wang X M, Zhang X, Zhang L W, Li Y, Huang G H. 2017. Evaluation on the responses of maize (Zea mays L.) growth, yield and water use efficiency to drip irrigation water under mulch condition in the Hetao irrigation district of China. Agricultural Water Management179, 144–157.

Liu H Q, Gao X D, Li C J, Cai Y H, Song X L, Zhao X N. 2025. Intercropping increases plant water availability and water use efficiency: A synthesis. AgricultureEcosystems and Environment379, 109360.

Liu L M, Klocke N, Yan S P, Rogers D, Schlegel A, Lamm F, Chang S I, Wang D H. 2013. Impact of deficit irrigation on maize physical and chemical properties and ethanol yield. Cereal Chemistry90, 453–462.

Liu R, Zhou G P, Li C J, Chang D N, Gao S J, Han M, Zhang J D, Sun X F, Cao W D. 2022. Transfer characteristics of nitrogen fixed by leguminous green manure crops when intercropped with maize in northwestern China. Journal of Integrative Agriculture21, 1177–1187.

Luo Y, Zhang J D, Zhou G P, Chang D N, Gao S J, Bao X G, Che Z X, Zhu Q, Cao W D. 2022. Intercropping maize with green manure crops at various utilization patterns improves maize yield and soil fertility in Hexi oasis irrigated area. Journal of Plant Nutrition and Fertilizer28, 402–413.

Lyu H Q, Li Y, Wang Y L, Wang P F, Shang Y P, Yang X H, Wang F, Yu A Z. 2023. Drive soil nitrogen transformation and improve crop nitrogen absorption and utilization - a review of green manure applications. Frontiers in Plant Science14, 15.

Ma Y N, Yang S J, Silva L, Ding R S, Du T S, Kang S Z, Tong L. 2025. Mild water deficit under reasonable dense planting ensures maize yield and improves water use efficiency by coordinating source-sink relationship. Field Crops Research327, 109888.

Mao L L, Zhang L Z, Li W Q, Van Der Werf W, Sun J H, Spiertz H, Li L. 2012. Yield advantage and water saving in maize/pea intercrop. Field Crops Research138, 11–20.

Marques G, Cavalcante K, Silva L, Brito N, Lima J, Moura E. 2018. Effects of leguminous green manure on the physicochemical quality of maize grains (Zea mays L.). Australian Journal of Crop Science12, 676–685.

Qiang B B, Fan Z, Tang N, Asad M S, Timbang B C, Ren X L, Chen X L. 2024. Improving the productivity of intercropping through above and below ground separation: A case study on photosynthetic characteristics and root distribution. Industrial Crops and Products222, 119506.

Shen S, Li B B, Deng T, Xiao Z D, Chen X M, Hu H, Zhang B C, Wu G, Li F, Zhao X, Liang X G, Mi G H, Zhou S L. 2020a. The equilibrium between sugars and ethylene is involved in shading- and drought-induced kernel abortion in maize. Plant Growth Regulation91, 101–111.

Shen S, Liang X G, Zhang L, Zhao X, Liu Y P, Lin S, Gao Z, Wang P, Wang Z M, Zhou S L. 2020b. Intervening in sibling competition for assimilates by controlled pollination prevents seed abortion under postpollination drought in maize. PlantCell and Environment43, 903–919.

Singh H, Northup B K, Prasad P V V. 2023. Water storage and use efficiencies of rainfed winter wheat-summer green manure systems of the US Southern Great Plains. European Journal of Agronomy146, 126818.

Tang H, Xu C S, Jiang Y M, Wang J W, Wang Z H, Tian L Q. 2021. Evaluation of physical characteristics of typical maize seeds in a cold area of North China based on principal component analysis. Processes9, 1167.

Tilman D. 2020. Benefits of intensive agricultural intercropping. Nature Plants6, 604–605.

Wang F, Wang Y L, Lyu H Q, Fan Z L, Hu F L, He W, Yin W, Zhao C, Chai Q, Yu A Z. 2023. No-tillage mulch with leguminous green manure retention reduces soil evaporation and increases yield and water productivity of maize. Agricultural Water Management290, 108573.

Wang W, Li M Y, Zhang W, Khan A, Zhou R, Zhu S G, Wang G Z, Yang Y M, Tao H Y, Li W B, Xiong Y C. 2024. Soil moisture drives the shift from selection to complementarity effect in the rainfed maize/faba bean intercropping system. Plant and Soil499, 313–328.

Wang X C, Deng X Y, Pu T, Song C, Yong T W, Yang F, Sun X, Liu W G, Yan Y H, Du J B, Liu J, Su K, Yang W Y. 2017. Contribution of interspecific interactions and phosphorus application to increasing soil phosphorus availability in relay intercropping systems. Field Crops Research, 204, 12–22.

Wang Y, Xu C X, Zhou G P, Chang D N, Gao S J, Cao W D. 2021. Effects of long-term incorporation of milk vetch combined with reduction of chemical fertilizer on yield, quality and soil fertility of early rice in Jiangxi. Journal of Plant Nutrition and Fertilizers27, 1735–1745.

Wu P, Zhao G, Huang H, Wu Q, Bangura K, Cai T, Sun M, Xue J F, Zhang J J, Dang Y, Wang S, Zhou G, Fu J, Yang K, Fan T, Gao Z. 2023. Optimizing soil and fertilizer management strategy to facilitate sustainable development of wheat production in a semi-arid area: A 12-year in-situ study on the Loess Plateau. Field Crops Research302, 109084.

Xia H Y, Qiao Y T, Li X J, Xue Y H, Wang N, Yan W, Xue Y F, Cui Z L, Werf W. 2023. Moderation of nitrogen input and integration of legumes via intercropping enable sustainable intensification of wheat-maize double cropping in the North China Plain: A four-year rotation study. Agricultural Systems204, 103540.

Xu Z, Li C J, Zhang C C, Yu Y, Van Der Werf W, Zhang F S. 2020. Intercropping maize and soybean increase efficiency of land and fertilizer nitrogen use: A meta-analysis. Field Crops Research246, 107661.

Yang R, Song S J, Chen S Y, Du Z Y, Kong J Q. 2023. Adaptive evaluation of green manure rotation for a low fertility farmland system: Impacts on crop yield, soil nutrients, and soil microbial community. Catena222, 106873.

Yao C S, Li J P, Zhang Z, Liu Y, Wang Z M, Sun Z C, Zhang Y H. 2023. Improving wheat yield, quality and resource utilization efficiency through nitrogen management based on micro-sprinkler irrigation. Agricultural Water Management282, 108277.

Yao Z Y, Zhang D B, Yao P W, Zhao N, Li Y Y, Zhang S Q, Zhai B N, Huang D L, Ma A S, Zuo Y J, Cao W D, Gao Y J. 2018. Optimizing the synthetic nitrogen rate to balance residual nitrate and crop yield in a leguminous green-manured wheat cropping system. Science of the Total Environment631–632, 1234–1242.

Yin W, Chai Q, Zhao C, Yu A Z, Fan Z L, Hu F L, Fan H, Guo Y, Coulter J A. 2020. Water utilization in intercropping: A review. Agricultural Water Management241, 106335.

Yin W, Yu A Z, Guo Y, Wang Y F, Zhao C, Fan Z L, Hu F L, Chai Q. 2018. Straw retention and plastic mulching enhance water use via synergistic regulation of water competition and compensation in wheat-maize intercropping systems. Field Crops Research229, 78–94.

Yu R, Zhang H Y, Chang F D, Song J S, Wang J, Wang X Q, Kan Z R, Zhao N, Li X H, Ma J, Li Y Y. 2024. Mixed sowing of feed rape and vicia villosa can substitute nitrogen fertilizer to improve soil multifunctionality in the Hetao irrigation district. Catena235, 107617.

Yu Y, Jiang Z H, Wang G J, Kattel G R, Chuai X W, Shang Y, Zou Y F, Miao L J. 2022. Disintegrating the impact of climate change on maize yield from human management practices in China. Agricultural and Forest Meteorology327, 109235.

Yue K, Li L L, Xie J H, Liu Y Q, Xie J H, Anwar S, Fudjoe S K. 2022. Nitrogen supply affects yield and grain filling of maize by regulating starch metabolizing enzyme activities and endogenous hormone contents. Frontiers in Plant Science12, 798119.

Zhang J D, Bao X G, Cao W D, Che Z X, Hu Z Q, Lu B L. 2013. Effect of intercropping green manure crops on maize yield and soil fertility. Soil and Fertilizer Sciences in China4, 43–47. (in Chinese)

Zhang L, Liang Z Y, He X M, Meng Q F, Hu Y C, Schmidhalter U, Zhang W, Zou C Q, Chen X P. 2020. Improving grain yield and protein concentration of maize (zea mays L.) simultaneously by appropriate hybrid selection and nitrogen management. Field Crops Research249, 107754.

Zhang M X, Bai R, Nan M, Ren W, Wang C M, Shabala S, Zhang J L. 2022. Evaluation of salt tolerance of oat cultivars and the mechanism of adaptation to salinity. Journal of Plant Physiology273, 153708.

Zhang W X, Li P, Yin W, Chen G P, Fan Z L, Hu F L, Fan H, He W. 2023. Effects of multiple green manure after wheat combined with different levels of nitrogen fertilization on wheat yield, grain quality, and nitrogen utilization. Scientia Agricultura Sinica56, 3317–3330. (in Chinese)

Zhang Y, Sun Z X, Su Z C, Du G J, Bai W, Wang Q, Wang R N, Nie J Y, Sun T R, Feng C, Zhang Z, Yang N, Zhang X, Evers J B, Van Der Werf W, Zhang L Z. 2022. Root plasticity and interspecific complementarity improve yields and water use efficiency of maize/soybean intercropping in a water-limited condition. Field Crops Research282, 108523.

Zheng B C, Zhou Y, Chen P, Zhang X N, Du Q, Yang H, Wang X C, Yang F, Xiao T, Li L, Yang W Y, Yong T W. 2022. Maize–leguminous intercropping promotes N uptake through changing the root spatial distribution, leguminous nodulation capacity, and soil N availability. Journal of Integrative Agriculture21, 1755–1771.

Zhou Q, Jiang D, Dai W B, Cao W D, Jing Q. 2004. Effects of varied carbon and nitrogen supply on starch and protein accumulation in grain during early filling of wheat. Australian Journal of Agricultural Research55, 975–979.

Zhu K L, Song C Y, Liu J T, Gong M B, Wang S J, Song X Y, Li J. 2021. Unravelling the mechanisms of improving wheat growth, yield, and grain quality under long-term corn straw return plus N fertilizer mode. Journal of Soil Science and Plant Nutrition21, 3428–3436.

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