Please wait a minute...
Journal of Integrative Agriculture  2026, Vol. 25 Issue (10): 3959-3970    DOI: 10.1016/j.jia.2026.06.037
Special Focus: Advancing China–Africa Agricultural Science and Technology Cooperation: Innovation and Pathways Advanced Online Publication | Current Issue | Archive | Adv Search |
Effects of intercropping on tea yield, quality, and soil properties: A meta-analysis

Jie Zhang1, Shaobo Zhang1, Qiang Hu1, Qingyu Hong1, Xinfeng Jiang2, Liping Zhang1, Lan Zhang1, Zhengzhen Li1, Shibei Ge1, Jianyu Fu1, Xin Li1, Peng Yan1#

1 Tea Research Institute, Chinese Academy of Agricultural Sciences, Hangzhou 311300, China

2 Jiangxi Province Cash Crops Research Institute/Jiangxi Provincial Key Laboratory of Plantation and High Valued Utilization of Specialty Fruit Tree and Tea, Nanchang 330006, China

 Highlights 
● A meta-analysis was performed to quantify intercropping effects on tea yield, quality, and soil properties.
● Intercropping significantly increased tea yield by 11.90%, which was mainly regulated by tea age, initial soil pH, and mean annual temperature.
● Tea quality responses were primarily driven by initial soil conditions, whereas soil improvements were more strongly controlled by climatic factors.
● Leguminous intercropping is superior to other types in improving the overall benefits of tea plantations.

Download:  PDF in ScienceDirect  
Export:  BibTeX | EndNote (RIS)      
摘要  

间作作为一种传统的、可持续的农业实践,有助于提高土壤肥力,减少侵蚀,促进植物生长和代谢。目前间作在茶园中被广泛采用,但间作对产量、质量和土壤的影响是高度依赖于环境的,随着物种选择、气候和场地特定条件的变化而变化。为定量研究间作对茶叶产量、质量和土壤的影响及其驱动因素,收集了41篇文章(157组试验数据),利用 Meta 分析方法系统量化间作对茶叶产量、品质及土壤性质的影响,明确不同管理、气候和土壤条件下的效应差异。结果表明,间作下茶叶产量显著提高11.9%,茶叶中游离氨基酸(AAs)含量升高19.08%,茶多酚(TPs)含量降低7.16%,酚氨比显著降低,有助于提高茶叶品质。土壤质量方面,间作可以提高土壤养分有效性,以及土壤pH。其中土壤初始有机质和pH条件对茶叶产量和品质结果有显著影响,而间作植物类型,年平均温度(MAT)和年平均降雨量(MAP)等气候因素与土壤质量的改善密切相关。综上,间作有利于提高茶叶产量和品质,同时改善茶园的土壤条件,但需要挑选合适的间作品种,并根据当地气候、土壤条件和种植园年龄进行定制管理,从而实现产量-品质-养分效率的协同提升。



Abstract  

Intercropping is a traditional and sustainable agricultural practice that enhances soil fertility, reduces erosion, and promotes plant growth and metabolism.  Despite widespread adoption in tea plantations, intercropping effects on yield, quality, and soil are highly context-dependent, varying with species choice, climate, and site-specific conditions.  Using 157 paired observations from 41 articles, we performed a meta-analysis to quantify how intercropping affects tea yield, quality, and soil, and to determine the primary modulating factors.  Across all included reports, intercropping significantly increased tea yield by 11.9%.  It also elevated the levels of free amino acids (AAs) by 19.08% and reduced the levels of tea polyphenols (TPs) by 7.16%, leading to a notable decrease in the TP/AA ratio, which is associated with tea quality.  Additionally, intercropping increased soil nutrient levels and soil pH.  Initial organic matter and pH had a significant influence on tea yield and quality outcomes, while the type of plants used for intercropping and climatic factors such as mean annual temperature and mean annual precipitation were closely associated with improvements in soil quality.  Overall, intercropping can improve tea yield and quality while enhancing soil conditions in tea plantations, although the magnitude of these benefits depends on companion crop selection, soil status, and climate.  

Keywords:  intercropping       tea plantation        quality        yield        soil quality  
Received: 16 January 2026   Accepted: 05 June 2026 Online: 27 June 2026  
Fund: This research was funded by the Key Research and Development Project of Science and Technology Department of Zhejiang Province, China (2023C02019), the Zhejiang Province Science and Technology Cooperation Project for Nine Affiliations of Agriculture, Rural Affairs, and Farmers, China (2024SNJF031), the Agricultural Science and Technology Innovation Program of Chinese Academy of Agricultural Sciences (CAAS-ZDRW202417), the Key Scientific Research Project of Tea Research Institute, Chinese Academy of Agricultural Sciences (TRI-ZDRW-01-04), the Modern Agricultural Industrial Technology System of Jiangxi Province, China (JXARS-06).
About author:  Jie Zhang, E-mail: zj19392024@163.com; #Correspondence Peng Yan, E-mail: yanpengzn@tricaas.com

Cite this article: 

Jie Zhang, Shaobo Zhang, Qiang Hu, Qingyu Hong, Xinfeng Jiang, Liping Zhang, Lan Zhang, Zhengzhen Li, Shibei Ge, Jianyu Fu, Xin Li, Peng Yan. 2026. Effects of intercropping on tea yield, quality, and soil properties: A meta-analysis. Journal of Integrative Agriculture, 25(10): 3959-3970.

Ahammed G J, Wu Y X, Wang Y M, Guo T M, Shamsy R, Li X. 2023. Epigallocatechin-3-gallate (EGCG): A unique secondary metabolite with diverse roles in plant–environment interaction. Environmental and Experimental Botany, 209, 105299.

Akanvou R, Bastiaans L, Kropff M J, Goudriaan J, Becker M. 2001. Characterization of growth, nitrogen accumulation and competitive ability of six tropical legumes for potential use in intercropping systems. Journal of Agronomy and Crop Science, 187, 111–120.

Azapagic A, Bore J, Cheserek B, Kamunya S, Elbehri A. 2016. The global warming potential of production and consumption of Kenyan tea. Journal of Cleaner Production, 112, 4031–4040.

Boonerjee S, Islam M A, Islam S M M, Paul S K, Uddin M T, Alamgir M S. 2025. Consumption and export potential of tea in bangladesh: A field study. Journal of Agriculture and Food Research, 19, 101607.

Brooker R W, Bennett A E, Cong W, Daniell T J, George T S, Hallett P D, Hawes C, Iannetta P P M, Jones H G, Karley A J, Li L, McKenzie B M, Pakeman R J, Paterson E, Schöb C, Shen J, Squire G, Watson C A, Zhang C, Zhang F, et al. 2015. Improving intercropping: A synthesis of research in agronomy, plant physiology and ecology. New Phytologist, 206, 107–117.

Cao R, Yang J C, Meng Z T, Zhou H, Dong X, Peng L, Hou S S. 2025. Linking microbial communities and organic matter dynamics in longjing and fuding tea ecosystems. Journal of Food Biochemistry, 2025, 9981444.

Chen D, Dou Q P. 2008. Tea polyphenols and their roles in cancer prevention and chemotherapy. International Journal of Molecular Sciences, 9, 1196–1206.

Chen M J, Zhou P Q, Bao Q, Wang H, Wang Y J, Fu H P. 2025. Intercropping different legumes in tea plantation improves soil properties and tea quality components by regulating rhizosphere soil microorganisms. Agronomy, 15, 511.

Chen Y, Li J W, Yang N, Hu Z H, Luo W, Chen C, Wang Y H, Chen X, Li X H, Zhuang J. 2025. Nitrogen forms and nitrogen deficiency regulate theanine accumulation patterns in tea plants (Camellia sinensis) during winter dormancy. Horticulturae, 11, 444.

Chen Y Y, Li J L, Zhou B, Wu X M, Cui Y Y, Feng S M, Hu H T, Tang J C. 2023. Effects of intercropping with Vulpia myuros in tea plantation on soil and tea quality components. Scientia Agricultura Sinica, 56, 4916–4929. (in Chinese)

Cong W F, Hoffland E, Li L, Janssen B H, van der Werf W. 2015. Intercropping affects the rate of decomposition of soil organic matter and root litter. Plant and Soil, 391, 399–411.

Dechassa N, Merga M. 2022. Tea (Camellia sinensis) production, opportunities, challenges, and future prospects in Ethiopia. Advances in Agriculture, 2022, 1942666.

Deng W W, Fei Y, Wang S, Wan X C, Zhang Z Z, Hu X Y. 2013. Effect of shade treatment on theanine biosynthesis in Camellia sinensis seedlings. Plant Growth Regulation, 71, 295–299.

Deng W W, Ogita S, Ashihara H. 2008. Biosynthesis of theanine (γ-ethylamino-l-glutamic acid) in seedlings of Camellia sinensis. Phytochemistry Letters, 1, 115–119.

Duan Y, Wang G, Liang L Y, Wang M H, Jiang J, Ma Y C, Zhu X J, Wu J, Fang W P. 2024a. Intercropping fruit trees in tea plantation improves soil properties and the formation of tea quality components. Plant Physiology and Biochemistry, 210, 108574.

Duan Y, Wang T, Lei X G, Cao Y, Liu L F, Zou Z W, Ma Y C, Zhu X J, Fang W P. 2024b. Leguminous green manure intercropping changes the soil microbial community and increases soil nutrients and key quality components of tea leaves. Horticulture Research, 11, uhae018.

Duan Y, Wang T, Zhang P X, Zhao X J, Jiang J, Ma Y C, Zhu X J, Fang W P. 2024c. The effect of intercropping leguminous green manure on theanine accumulation in the tea plant: A metagenomic analysis. Plant, Cell & Environment, 47, 1141–1159.

Fang C Y, Shen Y F, Wang F Y, Zhang J Y, Liu C, Luo F, Ye Y L. 2024. Combined transcriptomics and metabolomics to elucidate the underlying beneficial mechanisms of L-theanine in mitigating obesity-induced cardiac injury in rats. Journal of Functional Foods, 113, 106037.

Feng L, Yu Y C, Lin S J, Yang T Y, Chen Q, Liu L L, Sun J, Zheng P C, Zhang Z L, Wan X C. 2021. Tonoplast-localized theanine transporter CsCAT2 may mediate theanine storage in the root of tea plants (Camellia sinensis L.). Frontiers in Plant Science, 12, 797854.

Feng Y T, Sunderland T. 2023. Feasibility of tea/tree intercropping plantations on soil ecological service function in China. Agronomy, 13, 1548.

Fu X M, Cheng S H, Liao Y Y, Xu X L, Wang X C, Hao X Y, Xu P, Dong F, Yang Z Y. 2020. Characterization of L-theanine hydrolase in vitro and subcellular distribution of its specific product ethylamine in tea (Camellia sinensis). Journal of Agricultural and Food Chemistry, 68, 10842–10851.

Fung K F, Carr H P, Zhang J, Wong M H. 2008. Growth and nutrient uptake of tea under different aluminium concentrations. Journal of the Science of Food and Agriculture, 88, 1582–1591.

Godfray H C J, Poore J, Ritchie H. 2024. Opportunities to produce food from substantially less land. BMC Biology, 22, 138.

Hao T, Cui L J, Wang J Z, Lei Y R, Li W, Wang R M, Wang S K, Li J, Zhai X J, Zhang M Y, Zhao X S. 2025. Effects of low temperature on rhizosphere phosphate-mineralizing microbial populations in constructed wetlands. Journal of Environmental Management, 376, 124243.

Huang Z, Cui C H, Cao Y J, Dai J H, Cheng X Y, Hua S W, Wang W T, Duan Y, Petropoulos E, Wang H, Zhou L X, Fang W P, Zhong Z T. 2022. Tea plant–legume intercropping simultaneously improves soil fertility and tea quality by changing Bacillus species composition. Horticulture Research, 9, uhac046.

Hung Y, Chen P, Chen R L C, Cheng T. 2010. Sequential determination of tannin and total amino acid contents in tea for taste assessment by a fluorescent flow-injection analytical system. Food Chemistry, 118, 876–881.

Jia J Y, Zhang J Z, Li Y Z, Koziol L, Podzikowski L, Delgado-Baquerizo M, Wang G Z, Zhang J L. 2023. Relationships between soil biodiversity and multifunctionality in croplands depend on salinity and organic matter. Geoderma, 429, 116273.

Jiang Y H, Lin X Q, Wang H B, Xu Y N, Lin W X. 2025. Effect of intercropping with forage soybean on the rhizosphere soil nutrients of tea plants. Catena, 255, 109060.

Juneja L R, Chu D, Okubo T, Nagato Y, Yokogoshi H. 1999. L-theanine - a unique amino acid of green tea and its relaxation effect in humans. Trends in Food Science & Technology, 10, 199–204.

Kito M, Kokura H, Izaki J, Sasaoka K. 1968. Theanine, a precursor of the phloroglucinol nucleus of catechins in tea plants. Phytochemistry, 7, 599–603.

Lafleur B, Lalonde O, Labrecque M. 2017. First-rotation performance of five short-rotation willow cultivars on different soil types and along a large climate gradient. BioEnergy Research, 10, 158–166.

Lal R. 2009. Challenges and opportunities in soil organic matter research. European Journal of Soil Science, 60, 158–169.

Li N H, Gao D M, Zhou X G, Chen S C, Li C X, Wu F Z. 2020. Intercropping with potato-onion enhanced the soil microbial diversity of tomato. Microorganisms, 8, 834.

Li X F, Wang Z G, Bao X G, Sun J H, Yang S C, Wang P, Wang C B, Wu J P, Liu X R, Tian X L, Wang Yu, Li J P, Wang Y, Xia H Y, Mei P P, Wang X F, Zhao J H, Yu R P, Zhang W P, Che Z X, et al. 2021. Long-term increased grain yield and soil fertility from intercropping. Nature Sustainability, 4, 943–950.

Li X X, Zhang X H, Zhao Q S, Liao H. 2023. Genetic improvement of legume roots for adaption to acid soils. The Crop Journal, 11, 1022–1033.

Li Y H, Shi D Y, Li G H, Zhao B, Zhang J W, Liu P, Ren B Z, Dong S T. 2019. Maize/peanut intercropping increases photosynthetic characteristics, 13C-photosynthate distribution, and grain yield of summer maize. Journal of Integrative Agriculture, 18, 2219–2229.

Liu C G, Nie Y, Rao X, Tang J W, Siddique K H M. 2019. The effects of introducing Flemingia macrophylla to rubber plantations on soil water content and exchangeable cations. Catena, 172, 480–487.

Liu P F, Zhao Y K, Ma J N, Cao Y, Zhang M X, Yu J, Guan H B, Xing Y S, Wang X Q, Jia X. 2025. Impact of various intercropping modes on soil quality, microbial communities, yield and quality of Platycodon grandiflorum (Jacq.) A. DC. BMC Plant Biology, 25, 503.

Liu Y R, Lan X J, Hou H Q, Ji J H, Liu X M, Lv Z Z. 2024. Multifaceted ability of organic fertilizers to improve crop productivity and abiotic stress tolerance: Review and perspectives. Agronomy, 14, 1141.

Liu Z W, Wang Q, Wang T, Chen Q Y, He X F, Ma Y, Zhao M. 2025. Interplanting with leguminous plants improves theanine accumulation in tea shoots via the rhizosphere soil microecology. Industrial Crops and Products, 230, 121075.

Nowogrodzki A. 2019. How climate change might affect tea. Nature, 566, S10–S11.

Oldfield E E, Wood S A, Bradford M A. 2018. Direct effects of soil organic matter on productivity mirror those observed with organic amendments. Plant and Soil, 423, 363–373.

Osland M J, Gabler C A, Grace J B, Day R H, McCoy M L, McLeod J L, From A S, Enwright N M, Feher L C, Stagg C L, Hartley S B. 2018. Climate and plant controls on soil organic matter in coastal wetlands. Global Change Biology, 24, 5361–5379.

Ranjitkar S, Sujakhu N M, Lu Y, Wang Q, Wang M C, He J, Mortimer P E, Xu J C, Kindt R, Zomer R J. 2016. Climate modelling for agroforestry species selection in yunnan province, China. Environmental Modelling & Software, 75, 263–272.

Ruan J, Haerdter R, Gerendás J. 2010. Impact of nitrogen supply on carbon/nitrogen allocation: A case study on amino acids and catechins in green tea [Camellia sinensis (L.) O. Kuntze] plants. Plant Biology, 12, 724–734.

Ruan J Y, Ma L F, Yang Y J. 2012. Magnesium nutrition on accumulation and transport of amino acids in tea plants. Journal of the Science of Food and Agriculture, 92, 1375–1383.

Sahu N, Nayan R, Panda A, Varun A, Kesharwani R, Das P, Kumar A, Mallick S K, Mishra M M, Saini A, Aggarwal S P, Nayak S. 2025. Impact of changes in rainfall and temperature on production of Darjeeling tea in India. Atmosphere, 16, 1.

Shao S B, Li Z W, Ma X X, Cui J R, Zhu Y Q, Li Y P, Wu L K, Rensing C, Cai P M, Zhang J M, Li Q S. 2025. Enhancing tea plant growth and soil microbial ecology through intercropping tea plants with Ophiopogon japonicus. Plant and Soil, 513, 2807–2825.

Sun L T, Dong X, Wang Y, Maker G, Agarwal M, Ding Z T. 2022. Tea–soybean intercropping improves tea quality and nutrition uptake by inducing changes of rhizosphere bacterial communities. Microorganisms, 10, 2149.

Tilman D. 2020. Benefits of intensive agricultural intercropping. Nature Plants, 6, 604–605.

Tilman D, Balzer C, Hill J, Befort B L. 2011. Global food demand and the sustainable intensification of agriculture. Proceedings of the National Academy of Sciences of the United States of America, 108, 20260–20264.

Tilman D, Cassman K G, Matson P A, Naylor R, Polasky S. 2002. Agricultural sustainability and intensive production practices. Nature, 418, 671–677.

Vuong Q V, Bowyer M C, Roach P D. 2011. L-theanine: Properties, synthesis and isolation from tea. Journal of the Science of Food and Agriculture, 91, 1931–1939.

Wang D, Liu B J, Li F, Wang Z H, Hou J F, Cao R, Zheng Y Q, Yang W Q. 2025. Status and influential factors of soil nutrients and acidification in Chinese tea plantations: A meta-analysis. Soil, 11, 175–191.

Wang J F, Zhou Q, Lv Y L, Chen Z M. 2023. Effects of intercropping tea with landscape trees on ecosystem of tea garden and tea production. Acta Agriculturae Zhejiangensis, 35, 523–533. (in Chinese)

Wei H, Li H R, Wang Q, Xiang H M, Liu Z Q, Zhang J. 2025. Soil acidification alters C:N:P stoichiometry in the soil due to higher acid sensitivity of phosphorus. Environmental Science: Processes & Impacts, 27, 2094–2103.

Wen B, Zhang X L, Ren S, Duan Y, Zhang Y Y, Zhu X J, Wang Y H, Ma Y C, Fang W P. 2020. Characteristics of soil nutrients, heavy metals and tea quality in different intercropping patterns. Agroforestry Systems, 94, 963–974.

Wen X, Wu D L, Chen D J, Xu P R, Zhao T T, Chen S Y, Zhu Z H, Zhong H, Chen P. 2024. Soil organic matter and total nitrogen as key driving factors promoting the assessment of acid-base buffering characteristics in a tea (Camellia sinensis) plantation habitat. Environmental Monitoring and Assessment, 196, 596.

Wezel A, Casagrande M, Celette F, Vian J F, Ferrer A, Peigné J. 2014. Agroecological practices for sustainable agriculture. A review. Agronomy for Sustainable Development, 34, 1–20.

Wu H, Long X F, Geng Y F. 2023. Companion plants of tea: From ancient to terrace to forest. Plants, 12, 3061.

Wu Z H, Xue B, Wang S W, Xing X, Nuo M, Meng X, Wu M K, Jiang H, Ma H M, Yang M Y, Wei X S, Zhao G X, Tian P. 2024. Rice under dry cultivation–maize intercropping improves soil environment and increases total yield by regulating belowground root growth. Plants, 13, 2957.

Yan P, Wu L Q, Wang D H, Fu J Y, Shen C, Li X, Zhang L P, Zhang L, Fan L C, Han W Y. 2020. Soil acidification in Chinese tea plantations. Science of the Total Environment, 715, 136963.

Yan Z M, Zhong Y Z, Duan Y H, Chen Q H, Li F. 2020. Antioxidant mechanism of tea polyphenols and its impact on health benefits. Animal Nutrition, 6, 115–123.

Yang T Y, Xie Y X, Lu X, Yan X M, Wang Y, Ma J Z, Cheng X M, Lin S J, Bao S L, Wan X C, Lucas W J, Zhang Z L. 2021. Shading promoted theanine biosynthesis in the roots and allocation in the shoots of the tea plant (Camellia sinensis L.) cultivar shuchazao. Journal of Agricultural and Food Chemistry, 69, 4795–4803.

Yang X D, Ni K, Shi Y Z, Yi X Y, Zhang Q F, Fang L, Ma L F, Ruan J Y. 2018. Effects of long-term nitrogen application on soil acidification and solution chemistry of a tea plantation in China. Agriculture, Ecosystems & Environment, 252, 74–82.

Yang X D, Yi X Y, Ni K, Zhang Q F, Shi Y Z, Chen L B, Zhao Y Y, Zhang Y L, Ma Q X, Cai Y J, Ma L F, Ruan J Y. 2023. Patterns and abiotic drivers of soil organic carbon in perennial tea (Camellia sinensis L.) plantation system of China. Environmental Research, 237, 116925.

Ye J H, Wang H B, Yang X Y, Zhang Q, Li J Y, Jia X L, Kong X H, He H B. 2016. Autotoxicity of the soil of consecutively cultured tea plantations on tea (Camellia sinensis) seedlings. Acta Physiologiae Plantarum, 38, 195.

Ye J H, Wang Y H, Wang Y C, Hong L, Jia X L, Kang J Q, Lin S X, Wu Z Y, Wang H B. 2022. Improvement of soil acidification in tea plantations by long-term use of organic fertilizers and its effect on tea yield and quality. Frontiers in Plant Science, 13, 1055900.

Zhang J, Hu R B, Sreedasyam A, Garcia T M, Lipzen A, Wang M, Yerramsetty P, Liu D G, Ng V, Schmutz J, Cushman J C, Borland A M, Pasha A, Provart N J, Chen J G, Muchero W, Tuskan G A, Yang X H. 2020. Light-responsive expression atlas reveals the effects of light quality and intensity in Kalanchoë fedtschenkoi, a plant with crassulacean acid metabolism. GigaScience, 9, giaa018.

Zhang W J, Ni K, Long L Z, Ruan J Y. 2023. Nitrogen transport and assimilation in tea plant (Camellia sinensis): A review. Frontiers in Plant Science, 14, 1249202.

Zhao S X, Bai Y L, Jin Z, Long L, Diao W T, Chen W, Tan L Q, Tang Q, Tang D D. 2023. Effects of the combined application of nitrogen and selenium on tea quality and the expression of genes involved in nitrogen uptake and utilization in tea cultivar ‘Chuancha No.2.’ Agronomy, 13, 2997.

Zhao Y, Tian Y Q, Li X B, Song M H, Fang X X, Jiang Y, Xu X L. 2022. Nitrogen fixation and transfer between legumes and cereals under various cropping regimes. Rhizosphere, 22, 100546.

Zou Y, Shen F Y, Zhong Y N, Lv C N, Pokharel S S, Fang W P, Chen F J. 2022a. Impacts of intercropped maize ecological shading on tea foliar and functional components, insect pest diversity and soil microbes. Plants, 11, 1883.

Zou Y, Zhong Y N, Yu H, Pokharel S S, Fang W P, Chen F J. 2022b. Impacts of ecological shading by roadside trees on tea foliar nutritional and bioactive components, community diversity of insects and soil microbes in tea plantation. Biology, 11, 1800.

[1] Diaoliang Zhang, Yunyou Nan, Zhilong Fan, Qiang Chai, Gary Y. Gan, Wen Yin, Falong Hu. Optimizing maize yield and kernel quality via leguminous green manure intercropping with deficit irrigation in arid agroecosystem[J]. >Journal of Integrative Agriculture, 2026, 25(7): 3017-3030.
[2] Ping Lin, Shanshan Liu, Zhidan Fu, Kai Luo, Yiling Li, Xinyue Peng, Xiaoting Yuan, Lida Yang, Tian Pu, Yuze Li, Taiwen Yong, Wenyu Yang. Rhizosphere flavonoids alleviate inhibition of soybean nodulation caused by shading under maize–soybean strip intercropping[J]. >Journal of Integrative Agriculture, 2026, 25(3): 952-964.
[3] Chengzhi Jiao, Mingxing Wen, Xin Jing, Vanika Garg, Chuanqing Zhou, Liyang Chen, Fengfeng Xu, Chenyang Hao, Jin Xiao, Haiyan Wang, Rajeev K. Varshney, Xueyong Zhang, Xiu’e Wang. Accumulation of beneficial haplotypes in the Huang-Huai-Hai wheat region and its application in molecular breeding[J]. >Journal of Integrative Agriculture, 2026, 25(10): 4014-4026.
[4] Xin Zhao, Hai Liang, Danna Chang, Jiudong Zhang, Xingguo Bao, Heng Cui, Weidong Cao. Maize–green manure intercropping improves maize yield and P uptake by shaping the responses of roots and soil [J]. >Journal of Integrative Agriculture, 2026, 25(1): 313-325.
[5] Xiaohui Xu, Qiang Chai, Falong Hu, Wen Yin, Zhilong Fan, Hanting Li, Zhipeng Liu, Qiming Wang. Intercropping grain crops with green manure under reduced chemical nitrogen improves the soil carbon stocks by optimizing aggregates in an oasis irrigation area[J]. >Journal of Integrative Agriculture, 2026, 25(1): 326-338.
[6] Yang Chen, Xuyu Feng, Xiao Zhao, Xinmei Hao, Ling Tong, Sufen Wang, Risheng Ding, Shaozhong Kang. Biochar application enhances soil quality by improving soil physical structure under particular water and salt conditions in arid region of Northwest China[J]. >Journal of Integrative Agriculture, 2025, 24(8): 3242-3263.
[7] Xinhu Guo, Jinpeng Chu, Yifan Hua, Yuanjie Dong, Feina Zheng, Mingrong He, Xinglong Dai. Long-term integrated agronomic optimization maximizes soil quality and synergistically improves wheat yield and nitrogen use efficiency[J]. >Journal of Integrative Agriculture, 2025, 24(8): 2940-2953.
[8] Hanting Li, Zhilong Fan, Falong Hu, Wen Yin, Qiming Wang, Guocui Wang, Weidong Cao, Wei He, Qiang Chai, Tuo Yao. Intercropping maize with leguminous green manure can compensate for the losses in grain yield and N uptake caused by a reduced N supply[J]. >Journal of Integrative Agriculture, 2025, 24(7): 2826-2840.
[9] Liang Wang, Nijiang Ai, Zechang Zhang, Chenhui Zhou, Guoli Feng, Sheng Cai, Ningshan Wang, Liuchun Feng, Yu Chen, Min Xu, Yingying Wang, Haoran Yue, Mengfei Chen, Liangshuai Xing, Baoliang Zhou. Development of Gossypium hirsutum–Gossypium raimondii introgression lines and their use in QTL mapping of agricultural traits[J]. >Journal of Integrative Agriculture, 2025, 24(5): 1688-1703.
[10] Jia Wu, Luqi Zhang, Ziyi Wang, Fan Ge, Hao Zhang, Jianchang Yang, Yajie Zhang. Reasonable dry cultivation methods can balance the yield and grain quality of rice[J]. >Journal of Integrative Agriculture, 2025, 24(3): 1030-1043.
[11] Qingyun Tang, Guodong Wang, Lei Zhao, Zhiwen Song, Yuxiang Li.
Responses of yield, root traits and their plasticity to the nitrogen environment in nitrogen-efficient cultivars of drip-irrigated rice
[J]. >Journal of Integrative Agriculture, 2025, 24(2): 480-496.
[12] Yongshui Hao, Xueying Liu, Qianqian Wang, Shuxin Wang, Qingqing Li, Yaqing Wang, Zhongni Guo, Tiantian Wu, Qing Yang, Yuting Bai, Yuru Cui, Peng Yang, Wenwen Wang, Zhonghua Teng, Dexin Liu, Kai Guo, Dajun Liu, Jian Zhang, Zhengsheng Zhang. Mapping QTLs for fiber- and seed-related traits in Gossypium tomentosum CSSLs with a G. hirsutum background [J]. >Journal of Integrative Agriculture, 2025, 24(2): 467-479.
[13] Taowen Pan, Yulin Chen, Sicong Li, Lei Wang, Joji Muramoto, Carol Shennan, Jihui Tian, Kunzheng Cai. Anaerobic soil disinfestation rather than Bacillus velezensis Y6 inoculant suppresses tomato bacterial wilt by improving soil quality and manipulating bacterial communities[J]. >Journal of Integrative Agriculture, 2025, 24(2): 754-768.
[14] Xiaobo Gu, Zhikai Cheng, Yadan Du, Huanjie Cai, Yupeng Li, Yuannong Li, Heng Fang, Shikun Sun. Optimizing planting density to improve growth, yield and resource use efficiencies for winter oilseed rape under ridge-furrow film mulching[J]. >Journal of Integrative Agriculture, 2025, 24(10): 3819-3837.
[15] Fei Bao, Ping Zhang, Qiying Yu, Yunfei Cai, Bin Chen, Heping Tan, Hailiang Han, Junfeng Hou, Fucheng Zhao. Response of fresh maize yield to nitrogen application rates and  characteristics of nitrogen-efficient varieties[J]. >Journal of Integrative Agriculture, 2025, 24(10): 3803-3818.
No Suggested Reading articles found!