Adams S R, Cockshull K E, Cave C R J. 2001. Effect of temperature on the growth and development of tomato fruit. Ann Bot, 88(5), 869-877.

Albornoz K, Zhou J, Beckles DM. 2023. Chemical induction of the Arabidopsis thaliana CBF1 gene in transgenic tomato fruit to study postharvest chilling injury. Curr Plant Biol, 33, 100275.

Albornoz K, Zhou J, Zakharov F, Grove J, Wang M, Beckles D M. 2024. Ectopic overexpression of ShCBF1 and SlCBF1 in tomato suggests an alternative view of fruit responses to chilling stress postharvest. Front Plant Sci, 15, 1429321.

Archbold D D, Pomper K W. 2003. Ripening pawpaw fruit exhibit respiratory and ethylene climacterics. Postharvest Biol Technol, 30(1), 99-103.

Bai L, Wu Y, Lin H, Su W, Fan Z. 2025. MaERF9 and MaERF113 transcription factors involve in chilling injury development by regulating membrane lipid metabolism of postharvest banana fruit. Postharvest Biol Technol, 219, 113230.

Cai X, Chen Y, Wang Y, Shen Y, Yang J, Jia B, Sun X, Sun M. 2023. A comprehensive investigation of the regulatory roles of OsERF096, an AP2/ERF transcription factor, in rice cold stress response. Plant Cell Rep, 42(12), 2011-2022.

Cao K, Wei Y, Chen Y, Jiang S, Chen X, Wang X, Shao X. 2021. PpCBF6 is a low-temperature-sensitive transcription factor that binds the PpVIN2 promoter in peach fruit and regulates sucrose metabolism and chilling injury. Postharvest Biol Technol, 181, 111681.

Chen S, Xu L, Wang Y, Mao B, Zhang X, Song Q, Cui F, Ma Y, Dong J, Wang K, Bi H. 2025. RsWRKY40 coordinates the cold stress response by integrating RsSPS1-mediated sucrose accumulation and the CBF-dependent pathway in radish (Raphanus sativus L.). Mol Horticulture, 5(1), 14.

Cheng J, Wen S, Xiao S, Lu B, Ma M, Bie Z. 2018. Overexpression of the tonoplast sugar transporter CmTST2 in melon fruit increases sugar accumulation. J Exp Bot, 69(3), 511-523.

Cui M, Pham M D, Hwang H, Chun C. 2021. Flower development and fruit malformation in strawberries after short-term exposure to high or low temperature. Sci Hortic, 288, 110308.

Dai N, Cohen S, Portnoy V, Tzuri G, Harel-Beja R, Pompan-Lotan M, Carmi N, Zhang G, Diber, A, Pollock S, Karchi H. 2011. Metabolism of soluble sugars in developing melon fruit: a global transcriptional view of the metabolic transition to sucrose accumulation. Plant Mol Biol, 76, 1-18.

Dai N, Petreikov M, Portnoy V, Katzir N, Pharr D M, Schaffer A A. 2006. Cloning and expression analysis of a UDP-galactose/glucose pyrophosphorylase from melon fruit provides evidence for the major metabolic pathway of galactose metabolism in raffinose oligosaccharide metabolizing plants. Plant Physiol, 142(1), 294-304.

Fang T, Li Y, Xie T, Xian H, Bao Y, Zeng L. 2025. The bHLH transcription factor DlbHLH68 positively regulates DlSPS1 expression to promote sucrose biosynthesis in longan. Int J Biol Macromol, 296, 139594.

Fu M, Zheng Y, Zhang J, Deng C, Zhang J, Jia C, Miao H, Wang J, Zheng S, Jin Z, Li X. 2024. The MaEIL4-MaMADS36-MaACS7 module transcriptionally regulates ethylene biosynthesis during banana fruit ripening. Hortic Res, uhae345.

Galli F, Archbold D D, Pomper K W. 2008. Loss of ripening capacity of pawpaw fruit with extended cold storage. J Agric Food Chem, 56(22), 10683-10688.

Gao G, Yang F, Wang C, Duan X Y, Li M, Ma Y, Wang F, Qi H Y. 2023. The transcription factor CmERFI-2 represses CmMYB44 expression to increase sucrose levels in oriental melon fruit. Plant Physiol, 192(2), 1378-1395.

Gao J, Zhang Y, Li Z, Liu M. 2020. Role of ethylene response factors (ERFs) in fruit ripening. Food Qual Saf, m4(1), 15-20.

Gu J, Liu P, Nie W, Wang Z, Cui X, Fu H, Wang F, Qi M, Sun Z, Li T, Liu Y. 2025. Abscisic acid alleviates photosynthetic damage in the tomato ABA-deficient mutant sitiens and protects photosystem II from damage via the WRKY22–PsbA complex under low-temperature stress. J Integr Agric, 24(2), 546-563.

Gu S, Jing M, Li D, Ma Z, Duan Y, Wang L, Dai X, Chen Z, Zhang X, Chen J. 2024. The effects of different temperature and humidity conditions on the ripening and cracking of Annona atemoya fruit during storage by regulating the conversion of starch into soluble sugars. LWT, 208, 116703.

Guo C, Saren Q, Hao J, Guan X, Niu Y, Hasi A. 2019. In silico and expression profile analyses of the ERF subfamily in melon. Russ J Genet, 55(5), 557-570.

Guo N, Yan Y, Li Q, Yang Y. 2025. Pyruvic acid improves cold-storage quality of plum fruit by stimulating cyanide-resistant respiration and regulating sugar metabolism. Sci Hortic, 340, 113926.

Guo T, Li J, Guo M, Yang Q, Dai X, Qiao X, Song Z, Tian C, Li Y, Ge H, Cheng J. 2023. Low temperature inhibits pectin degradation by PpCBFs to prolong peach storage time. J Food Sci, 88(9), 3725-3736.

Hao J H, Yang R, Fang K F, Wang J L, Zhang Q, Shen Y Y, Li T L. 2014. Low night temperature inhibit galactinol synthase gene expression and phloem loading in melon leaves during fruit development. Russ J Plant Physiol, 61, 178-187.

Iraqi D, Tremblay FM. 2001. Analysis of carbohydrate metabolism enzymes and cellular contents of sugars and proteins during spruce somatic embryogenesis suggests a regulatory role of exogenous sucrose in embryo development. J Exp Bot, 52(365), 2301-2311.

Jia X, Wang W, Du Y, Wei T, Wang Z, Gul H. 2018. Optimal storage temperature and 1-MCP treatment combinations for different marketing times of Korla Xiang pears. J Integr Agric, 17(3), 693-703.

Kano Y. 2006. Effect of heating fruit on cell size and sugar accumulation in melon fruit (Cucumis melo L.). Hort Sci, 41(6), 1431-1434.

Khan M, Hu J, Dahro B, Ming R, Zhang Y, Wang Y, Alhag A, Li C, Liu J H. 2021. ERF108 from Poncirus trifoliata (L.) Raf. functions in cold tolerance by modulating raffinose synthesis through transcriptional regulation of PtrRafS. Plant J, 108(3), 705-724.

Lang X, Zhao X, Zhao J, Ren T, Nie L, Zhao W. 2024. MicroRNA Profiling Revealed the Mechanism of Enhanced Cold Resistance by Grafting in Melon (Cucumis melo L.). Plants, 13(7), 1016.

Li A, Zhang Z, Wang X C, Huang R. 2009. Ethylene response factor TERF1 enhances glucose sensitivity in tobacco through activating the expression of sugar‐related genes. J Integr Plant Biol, 51(2), 184-193.

Li B, Qu S, Kang J, Peng Y, Yang N, Ma B, Ruan Y L, Ma F, Li M, Zhu L. 2024. The MdCBF1/2‐MdTST1/2 module regulates sugar accumulation in response to low temperature in apple. Plant J, 118(3), 787-801.

Li L, Li Q, Chen B, Wang J, Ding F, Wang P, Zhang X, Hou J, Luo R, Li X, Zheng J. 2023. Identification of candidate genes that regulate the trade-off between seedling cold tolerance and fruit quality in melon (Cucumis melo L.). Hortic Res, 10(7), uhad093.

Li M, Duan X, Gao G, Liu T, Qi H. 2022. CmABF1 and CmCBF4 cooperatively regulate putrescine synthesis to improve cold tolerance of melon seedlings. Hortic Res, 9, uhac002.

Li S, Wu P, Yu X, Cao J, Chen X, Gao L, Chen K, Grierson D. 2022. Contrasting roles of ethylene response factors in pathogen response and ripening in fleshy fruit. Cells, 11(16), 2484.

Li X, Guo W, Li J, Yue P, Bu H, Jiang J, Liu W, Xu Y, Yuan H, Li T, Wang A. 2020. Histone acetylation at the promoter for the transcription factor PuWRKY31 affects sucrose accumulation in pear fruit. Plant Physiol, 182(4), 2035-2046.

Liu M, Pirrello J, Chervin C, Roustan J P, Bouzayen M. 2015. Ethylene control of fruit ripening: revisiting the complex network of transcriptional regulation. Plant Physiol, 169(4), 2380-2390.

Liu Y, He D, Wu Y, Zhao K, Yang C, Zhong Y, Yang L, Niu H, Liu S. 2025. Identification and Analysis of Melon (Cucumis melo L.) SHMT Gene Family Members and Their Functional Studies on Tolerance to Low-Temperature Stress. Agronomy, 15(1), 203.

Magnani E, Sjölander K, Hake S. 2004. From endonucleases to transcription factors: evolution of the AP2 DNA binding domain in plants. Plant Cell, 16(9), 2265-2277.

Mao W, Han Y, Chen Y, Sun M, Feng Q, Li L, Liu L, Zhang K, Wei L, Han Z, Li B. 2022. Low temperature inhibits anthocyanin accumulation in strawberry fruit by activating FvMAPK3-induced phosphorylation of FvMYB10 and degradation of Chalcone Synthase 1. Plant Cell, 34(4), 1226-1249.

Mata C I, Hertog M L, Van Raemdonck G, Baggerman G, Tran D, Nicolai B M. 2019. Omics analysis of the ethylene signal transduction in tomato as a function of storage temperature. Postharvest Biol Technol, 155, 1-10.

Musacchi S, Serra S. 2018. Apple fruit quality: Overview on pre-harvest factors. Sci Hortic, 234, 409-430.

Nakano T, Suzuki K, Fujimura T, Shinshi H. 2006. Genome-wide analysis of the ERF gene family in Arabidopsis and rice. Plant Physiol, 140(2), .411-432.

Pardossi A, Giacomet P, Malorgio F, Albini F M, Murelli C, Serra G, Vernieri P. 2000. The influence of growing season on fruit yield and quality of greenhouse melon (Cucumis melo L.) grown in nutrient film technique in a Mediterranean climate. J Hort Sci Biotechn, 75(4), 488–493. .

Paris H S, Amar Z, Lev E. 2012. Medieval emergence of sweet melons, Cucumis melo (Cucurbitaceae). Ann Bot, 110(1), 23-33. https://doi.org/10.1093/aob/mcs098.

Pech J C, Bouzayen M, Latché A J P S. 2008. Climacteric fruit ripening: ethylene-dependent and independent regulation of ripening pathways in melon fruit. Plant Sci, 175(1-2), 114-120.

Pujol M and Garcia-Mas J. 2023. Regulation of climacteric fruit ripening in melon: recent advances and future challenges. J Exp Bot, 74(20), 6224-6236.

Ren Y, Liao S, Xu Y. 2023. An update on sugar allocation and accumulation in fruit. Plant Physiol, 193(2), 888-899.

Roy Choudhury S, Roy S, Das R, Sengupta D N. 2008. Differential transcriptional regulation of banana sucrose phosphate synthase gene in response to ethylene, auxin, wounding, low temperature and different photoperiods during fruit ripening and functional analysis of banana SPS gene promoter. Planta, 229, 207-223.

Shi Y, Tian S, Hou L, Huang X, Zhang X, Guo H, Yan, S. 2012. Ethylene signaling negatively regulates freezing tolerance by repressing expression of CBF and type-A ARR genes in Arabidopsis. Plant Cell, 24(6), 2578-2595.

Tacken E, Ireland H, Gunaseelan K, Karunairetnam S, Wang D, Schultz K, Bowen J, Atkinson R G, Johnston J W, Putterill J, Hellens R P. 2010. The role of ethylene and cold temperature in the regulation of the apple POLYGALACTURONASE1 gene and fruit softening. Plant Physiol, 153(1), 294-305.

Tian H, Ma L, Zhao C, Hao H, Gong B, Yu X, Wang X. 2010. Antisense repression of sucrose phosphate synthase in transgenic muskmelon alters plant growth and fruit development. Biochem bioph res co, 393(3), 365-370.

Wang K L C, Li H, Ecker J R. 2002. Ethylene biosynthesis and signaling networks. Plant Cell, 14 (suppl_1), S131-S151.

Wang S Y, Camp M J. 2000. temperature after bloom affect plant growth and fruit quality of strawberry. Sci Hortic, 85(3), 183-199.

Wu X, Yu M, Huan C, Ma R, Yu Z. 2018. Regulation of the protein and gene expressions of ethylene biosynthesis enzymes under different temperature during peach fruit ripening. Acta Physiol Plant, 40, 1-9.

Xiao G, Qin H, Zhou J, Quan R, Lu X, Huang R, Zhang H. 2016. OsERF2 controls rice root growth and hormone responses through tuning expression of key genes involved in hormone signaling and sucrose metabolism. Plant Mol Biol, 90, 293-302.

Xiao X M, Si J, Wei W, Yang Y Y, Shan W, Kuang J F, Lu W J, Chen J Y, Chen J W. 2023. Banana ERF transcription factor MaERF110 is involved in ethylene-induced chilling tolerance by regulating ROS accumulation. Postharvest Biol Technol, 197, 112218.

Xu P Y, Xu L, Xu H F, He X W, He P, Chang Y S, Wang S, Zheng W Y, Wang C Z, Chen X, Li L G. 2023. MdWRKY40 is directly promotes anthocyanin accumulation and blocks MdMYB15L, the repressor of MdCBF2, which improves cold tolerance in apple. J Integr Agr, 22(6), 1704-1719.

Yang F, Gao G, Wang C, Guan J, Chen J, Qi H. 2025. CmERFⅠ-5 and CmERFⅤ-2 affect ethylene synthesis and sucrose accumulation in postharvest oriental melon fruit at low temperature. Postharvest Biol Technol, 219, 113295.

Yamada H, Hamamoto K, Sugiura A, Tomana T. 1988. Effect of controlled fruit temperature on maturation of apple fruit. J Jpn Soc Hortic Sci, 57(2), 173-177.

Yu W, Ma P, Sheng J, Shen L. 2023. Postharvest fruit quality of tomatoes influenced by an ethylene signaling component during long-term cold storage. Food Chem, 422, 136087.

Yu W, Shen L, Sheng J. 2024. Synergistic effects of SlCBF1 and ethylene signaling on the maintenance of tomatoes quality during long-term cold storage. Postharvest Biol Technol, 217, 113090.

Yu W, Sheng J, Zhao R, Wang Q, Ma P, Shen L. 2019. Ethylene biosynthesis is involved in regulating chilling tolerance and SlCBF1 gene expression in tomato fruit. Postharvest Biol Technol, 149, 139-147.

Yun Z E, Jin S, Ding Y, Wang Z, Gao H, Pan Z, Xu J, Cheng Y, Deng X. 2012. Comparative transcriptomics and proteomics analysis of citrus fruit, to improve understanding of the effect of low temperature on maintaining fruit quality during lengthy post-harvest storage. J Exp Bot, 63(8), 2873-2893.

Zhang B, Xi W P, Wei W W, Shen J Y, Ferguson I, Chen K S. 2011. Changes in aroma-related volatiles and gene expression during low temperature storage and subsequent shelf-life of peach fruit. Postharvest Biol Technol, 60(1), 7-16.

Zhang J, Yin X R, Li H, Xu M, Zhang M X, Li S J, Liu X F, Shi Y N, Grierson D, Chen K S. 2020. ETHYLENE RESPONSE FACTOR39–MYB8 complex regulates low-temperature-induced lignification of loquat fruit. J Exp Bot, 71(10), 3172-3184.

Zhang T, Zhang Q, Pan Y, Che F, Wang Q, Meng X, Rao J. 2017. Changes of polyamines and CBFs expressions of two Hami melon (Cucumis melo L.) cultivars during low temperature storage. Sci Hortic, 224, 8-16.

Zhang Z, Qi L, Zang N, Yang Y, Wang B, Liu X, Yin Z, Wang A. 2025. Jasmonate activated PuCBF5 to increasing ester accumulation in cold-stored ‘Nanguo’pear fruit. Plant Physiol Bioch, 109722.

Zhao D, Shen L, Fan B, Yu M, Zheng Y, Lv S, Sheng J. 2009. Ethylene and cold participate in the regulation of LeCBF1 gene expression in postharvest tomato fruit. FEBS Lett, 583(20), 3329-3334.

Zhu Y, Wu C, Wei W, Shan W, Kuang J, Chen J, Zhou E, Lu W, Yang Y. 2024. MaWRKY147-MaMADS68 transcriptional cascade module regulates low-temperature-affected banana fruit ripening. Postharvest Biol Technol, 207, 112625.