An J P, Li H H, Song LQ, Su L, Liu X, You C X, Wang X F, Hao Y J. 2016. The molecular cloning and functional characterization of MdMYC2, a bHLH transcription factor in apple. Plant Physiology and Biochemistry, 108, 24–31.

An J P, Wang X F, Zhang X W, You C X, Hao Y J. 2021. Apple BT2 protein negatively regulates jasmonic acid-triggered leaf senescence by modulating the stability of MYC2 and JAZ2. Plant, Cell & Environment, 44, 216–233.

Baker R F, Leach K A, Boyer N R, Swyers M J, Benitez-Alfonso Y, Skopelitis T, Luo A, Sylvester A, Jackson D, Braun D M. 2016. Sucrose transporter ZmSUT1 expression and localization uncover new insights into sucrose phloem loading. Plant Physiology, 172, 1876–1898.

Barth I, Meyer S, Sauer N. 2003. PmSUC3: characterization of a SUT2/SUC3-type sucrose transporter from Plantago major. Plant Cell, 15, 1375–1385.

Batistic O, Sorek N, Schultke S, Yalovsky S, Kudla J. 2008. Dual fatty acyl modification determines the localization and plasma membrane targeting of CBL/CIPK Ca2+ signaling complexes in Arabidopsis. Plant Cell, 20, 1346–1362.

Bihmidine S, Hunter C T, Johns C E, Koch K E, Braun D M. 2013. Regulation of assimilate import into sink organs: update on molecular drivers of sink strength. Frontiers in Plant Science, 4, 177.

Braun D M. 2022. Phloem loading and unloading of sucrose: What a long, strange trip from source to sink. Annual Review of Plant Biology, 73, 553–584.

Buttner M. 2010. The Arabidopsis sugar transporter (AtSTP) family: an update. Plant Biology, 12(Suppl. 1), 35–41.

Carpaneto A, Geiger D, Bamberg E, Sauer N, Fromm J, Hedrich R. 2005. Phloem-localized, proton-coupled sucrose carrier ZmSUT1 mediates sucrose efflux under the control of the sucrose gradient and the proton motive force. Journal of Biological Chemistry, 280, 21437–21443.

Chandran D, Reinders A, Ward J M. 2003. Substrate specificity of the Arabidopsis thaliana sucrose transporter AtSUC2. Journal of Biological Chemistry, 278, 44320–44325

Cheng J T, Wen S Y, Xiao S, Lu B Y, Ma M R, Bie Z L. 2018. Overexpression of the tonoplast sugar transporter CmTST2 in melon fruit increases sugar accumulation. Journal of Experimental Botany, 69, 511–523.

Chincinska I, Gier K, Krügel U, Liesche J, He H, Grimm B, Harren F, Cristescu S M, Kühn C. 2013. Photoperiodic regulation of the sucrose transporter StSUT4 affects the expression of circadian-regulated genes and ethylene production. Frontiers in Plant Science, 4, 26.

Chincinska I A, Liesche J, Krugel U, Michalska J, Geigenberger P, Grimm B, Kuhn C. 2008. Sucrose transporter StSUT4 from potato affects flowering, tuberization, and shade avoidance response. Plant Physiology, 146, 515–528.

Dhungana S R, Braun D M. 2021. Sugar transporters in grasses: Function and modulation in source and storage tissues. Journal of Plant Physiology, 266, 153541.

Dominguez P G, Niittyla T. 2022. Mobile forms of carbon in trees: metabolism and transport. Tree Physiology, 42, 458–487.

Durand M, Mainson D, Porcheron B, Maurousset L, Lemoine R, Pourtau N. 2018. Carbon source-sink relationship in Arabidopsis thaliana: the role of sucrose transporters. Planta, 247, 587–611.

Endler A, Meyer S, Schelbert S, Schneider T, Weschke W, Peters SW, Keller F, Baginsky S, Martinoia E, Schmidt U G. 2006. Identification of a vacuolar sucrose transporter in barley and Arabidopsis mesophyll cells by a tonoplast proteomic approach. Plant Physiology, 141, 196–207.

Feas X, Estevinho L M, Salinero C, Vela P, Sainz M J, Vazquez-Tato M P, Seijas J A. 2013. Triacylglyceride, antioxidant and antimicrobial features of virgin Camellia oleifera, C. reticulata and C. sasanqua oils. Molecules, 18, 4573–4587.

Frost C J, Nyamdari B, Tsai C J, Harding S A. 2012. The tonoplast-localized sucrose transporter in Populus (PtaSUT4) regulates whole-plant water relations, responses to water stress, and photosynthesis. PLoS One, 7, e44467.

Gong X, Liu M, Zhang L, Ruan Y, Ding R, Ji Y, Zhang N, Zhang S, Farmer J, Wang C. 2015. Arabidopsis AtSUC2 and AtSUC4, encoding sucrose transporters, are required for abiotic stress tolerance in an ABA-dependent pathway. Physiologia Plantarum, 153, 119–136.

Gottwald J R, Krysan P J, Young J C, Evert R F, Sussman M R. 2000. Genetic evidence for the in planta role of phloem-specific plasma membrane sucrose transporters. Proceedings of the National Academy of Sciences of the United States of America, 97, 13979–13984.

Hackel A, Schauer N, Carrari F, Fernie A R, Grimm B, Kuhn C. 2006. Sucrose transporter LeSUT1 and LeSUT2 inhibition affects tomato fruit development in different ways. Plant Journal, 45, 180–192.

Xu H F, Qu C Z, Liu J X, Wang Y C, Wang D Y, Zuo W F, Jiang S H, Wang N, Zhang Z Y, Chen X S. 2017. Expression analysis and functional identification of a vacuolar sucrose transporter gene MdSUT4 in apple. Acta Horticulturae Sinica, 44, 1235–1243. (in Chinese)

Hu Z, Tang Z, Zhang Y, Niu L, Yang F, Zhang D, Hu Y. 2021. Rice SUT and SWEET transporters. International Journal of Molecular Sciences, 22, 11198.

Javelle M, Timmermans M C. 2012. In situ localization of small RNAs in plants by using LNA probes. Nature Protocol, 7, 533–541.

Jia W Q, Zhang L J, Wu D, Liu S, Gong X, Cui Z H, Cui N, Cao H Y, Rao L B, Wang C. 2015. Sucrose transporter AtSUC9 mediated by a low sucrose level is involved in Arabidopsis abiotic stress resistance by regulating sucrose distribution and ABA accumulation. Plant & Cell Physiology, 56, 1574–1587.

Jiang M, Zhang J. 2001. Effect of abscisic acid on active oxygen species, antioxidative defence system and oxidative damage in leaves of maize seedlings. Plant & Cell Physiology, 42, 1265–1273.

Kariya K, Sameeullah M, Sasaki T, Yamamoto Y. 2017. Overexpression of the sucrose transporter gene NtSUT1 alleviates aluminum-induced inhibition of root elongation in tobacco (Nicotiana tabacum L.). Soil Science and Plant Nutrition, 63, 45–54.

Leach K A, Tran T M, Slewinski T L, Meeley R B, Braun D M. 2017. Sucrose transporter2 contributes to maize growth, development, and crop yield. Journal of Integrative Plant Biology, 59, 390–408.

Li M Z, Wang G P, Wu Y, Ren Y, Li G H, Liu Z H, Ding Y F, Chen L. 2020. Function analysis of sucrose transporter OsSUT4 in sucrose transport in rice. Chinese Journal of Rice Science, 34, 8.

Liang Y, Bai J, Xie Z, Lian Z, Guo J, Zhao F, Liang Y, Huo H, Gong H. 2023. Tomato sucrose transporter SlSUT4 participates in flowering regulation by modulating gibberellin biosynthesis. Plant Physiology, 192, 1080–1098.

Lin P, Wang K L, Wang Y P, Hu Z K, Yan C, Huang H, Ma X J, Cao Y Q, Long W, Liu W X, Li X L, Fan Z Q, Li J Y, Ye N, Ren H D, Yao X H, Yin H F. 2022. The genome of oil-Camellia and population genomics analysis provide insights into seed oil domestication. Genome Biology, 23, 14.

Lu M, Snyder R, Grant J, Tegeder M. 2020. Manipulation of sucrose phloem and embryo loading affects pea leaf metabolism, carbon and nitrogen partitioning to sinks as well as seed storage pools. Plant Journal, 101, 217–236.

Luo D, Xiong C, Lin A, Zhang C, Sun W, Zhang J, Yang C, Lu Y, Li H, Ye Z, He P, Wang T. 2021. SlBBX20 interacts with the COP9 signalosome subunit SlCSN5-2 to regulate anthocyanin biosynthesis by activating SlDFR expression in tomato. Horticulture Research, 8, 163.

Ma Q J, Sun M H, Liu Y J, Lu J, Hu D G, Hao Y J. 2016. Molecular cloning and functional characterization of the apple sucrose transporter gene MdSUT2. Plant Physiology and Biochemistry, 109, 442–451.

Meyer S, Melzer M, Truernit E, Hümmer C, Besenbeck R, Stadler R, Sauer N. 2000. AtSUC3, a gene encoding a new Arabidopsis sucrose transporter, is expressed in cells adjacent to the vascular tissue and in a carpel cell layer. Plant Journal, 24, 869–882.

Milne R J, Grof C P, Patrick J W. 2018. Mechanisms of phloem unloading: shaped by cellular pathways, their conductances and sink function. Current Opinion in Plant Biology, 43, 8–15.

Nebolisa N M, Umeyor C E, Ekpunobi U E, Umeyor I C, Okoye F B. 2023. Profiling the effects of microwave-assisted and soxhlet extraction techniques on the physicochemical attributes of Moringa oleifera seed oil and proteins. Oil Crop Science, 8, 16–26.

Patrick J W. 1997. Phloem unloading: Sieve element unloading and post-sieve element transport. Annual Review of Plant Physiology and Plant Molecular Biology, 48, 191–222.

Payyavula R S, Tay K H C, Tsai C J, Harding S A. 2011. The sucrose transporter family in Populus: the importance of a tonoplast PtaSUT4 to biomass and carbon partitioning. Plant Journal, 65, 757–770.

Pegler J L, Grof C P, Patrick J W. 2023. Sugar loading of crop seeds - a partnership of phloem, plasmodesmal and membrane transport. New Phytologist, 239, 1584–1602.

Peng Q, Cai Y, Lai E, Nakamura M, Liao L, Zheng B, Ogutu C, Cherono S, Han Y. 2020. The sucrose transporter MdSUT4.1 participates in the regulation of fruit sugar accumulation in apple. BMC Plant Biology, 20, 191.

Pugh D A, Offler C E, Talbot M J, Ruan Y L. 2010. Evidence for the role of transfer cells in the evolutionary increase in seed and fiber biomass yield in cotton. Molecular Plant, 3, 1075–1086

Radchuk V, Riewe D, Peukert M, Matros A, Strickert M, Radchuk R, Weier D, Steinbiss H H, Sreenivasulu N, Weschke W, Weber H. 2017. Down-regulation of the sucrose transporters HvSUT1 and HvSUT2 affects sucrose homeostasis along its delivery path in barley grains. Journal of Experimental Botany, 68, 4595–4612.

Raghavendra A S, Gonugunta V K, Christmann A, Grill E. 2010. ABA perception and signalling. Trends in Plant Science, 15, 395–401.

Reddy V S, Shlykov M A, Castillo R, Sun E I, Saier M H Jr. 2012. The major facilitator superfamily (MFS) revisited. FEBS Journal, 279, 2022–2035.

Reinders A, Sivitz A B, Starker C G, Gantt J S, Ward J M. 2008. Functional analysis of LjSUT4, a vacuolar sucrose transporter from Lotus japonicus. Plant Molecular Biology, 68, 289–299.

Ruan Y L. 2014. Sucrose metabolism: gateway to diverse carbon use and sugar signaling. Annual Review of Plant Biology, 65, 33–67.

Ruan Y L. 2022. CWIN-sugar transporter nexus is a key component for reproductive success. Journal of Plant Physiology, 268, 153572.

Ruan Y L, Llewellyn D J, Furbank R T. 2001. The control of single-celled cotton fiber elongation by developmentally reversible gating of plasmodesmata and coordinated expression of sucrose and K+ transporters and expansin. Plant Cell, 13, 47–60.

Sauer N. 2007. Molecular physiology of higher plant sucrose transporters. FEBS Letters, 581, 2309–2317.

Shah S T, Pang C, Fan S, Song M, Arain S, Yu S. 2013. Isolation and expression profiling of GhNAC transcription factor genes in cotton (Gossypium hirsutum L.) during leaf senescence and in response to stresses. Gene, 531, 220–234.

Shen S, Ma S, Chen X M, Yi F, Li B B, Liang X G, Liao S J, Gao L H, Zhou S L, Ruan Y L. 2022. A transcriptional landscape underlying sugar import for grain set in maize. Plant Journal, 110, 228–242.

Singh J, Thakur J K. 2018. Photosynthesis and abiotic stress in plants. In: S Vats, ed. Biotic and Abiotic Stress Tolerance in Plants; Springer: Cham, Switzerland. pp. 27–46.

Sosso D, Luo D, Li Q B, Sasse J, Yang J, Gendrot G, Suzuki M, Koch K E, McCarty D R, Chourey P S, Rogowsky P M, Ross-Ibarra J, Yang B, Frommer W B. 2015. Seed filling in domesticated maize and rice depends on SWEET-mediated hexose transport. Nature Genetics, 47, 1489–1493.

Srivastava A C, Ganesan S, Ismail I O, Ayre B G. 2008. Functional characterization of the Arabidopsis AtSUC2 sucrose/H+ symporter by tissue-specific complementation reveals an essential role in phloem loading but not in long-distance transport. Plant Physiology, 148, 200–211.

Sussmilch F C, Atallah N M, Brodribb T J, Banks J A, Mcadam S A M. 2017. Abscisic acid (ABA) and key proteins in its perception and signaling pathways are ancient, but their roles have changed through time. Plant Signaling & Behavior, 12, e1365210.

Thorne J H. 1985. Phloem unloading of C and N assimilates in developing seeds. Annual Review of Plant Physiology, 36, 317–343.

Wang D, Liu H, Wang H, Zhang P, Shi C. 2020. A novel sucrose transporter gene IbSUT4 involves in plant growth and response to abiotic stress through the ABF-dependent ABA signaling pathway in sweetpotato. BMC Plant Biology, 20, 157.

Wang L, Lu Q, Wen X, Lu C. 2015. Enhanced sucrose loading improves rice yield by increasing grain size. Plant Physiology, 169, 2848–2862.

Wang L F, Qi X X, Huang X S, Xu L L, Jin C, Wu J, Zhang S L. 2016. Overexpression of sucrose transporter gene PbSUT2 from Pyrus bretschneideri, enhances sucrose content in Solanum lycopersicum fruit. Plant Physiology and Biochemistry, 105, 150–161.

Wang X, You H, Yuan Y, Zhang H, Zhang L. 2018. The cellular pathway and enzymatic activity for phloem-unloading transition in developing Camellia oleifera Abel. fruit. Acta Physiologiae Plantarum, 40, 23.

Wang X Q, Zeng Q M, Contreras M D, Wang L J. 2017. Profiling and quantification of phenolic compounds in Camellia seed oils: Natural tea polyphenols in vegetable oil. Food Research International, 102, 184–194.

Wang Z, Wei X, Yang J, Li H, Ma B, Zhang K, Zhang Y, Cheng L, Ma F, Li M. 2020. Heterologous expression of the apple hexose transporter MdHT2.2 altered sugar concentration with increasing cell wall invertase activity in tomato fruit. Plant Biotechnology Journal, 18, 540–552.

Weschke W, Panitz R, Sauer N, Wang Q, Neubohn B, Weber H, Wobus U. 2000. Sucrose transport into barley seeds: molecular characterization of two transporters and implications for seed development and starch accumulation. Plant Journal, 21, 455–467.

Wieczorke R, Krampe S, Weierstall T, Freidel K, Hollenberg C P, Boles E. 1999. Concurrent knock-out of at least 20 transporter genes is required to block uptake of hexoses in Saccharomyces cerevisiae. FEBS Letters, 464, 123–128.

Yadav U P, Evers J F, Shaikh M A, Ayre B G. 2022. Cotton phloem loads from the apoplast using a single member of its nine-member sucrose transporter gene family. Journal of Experimental Botany, 73, 848–859.

Yan N. 2013. Structural advances for the major facilitator superfamily (MFS) transporters. Trends in Biochemical Sciences, 38, 151–159.

Yang S, Liang K, Sun Y, Zhang J, Cao Y, Zhou J, Wang A, Zhang L. 2023. Evidence of the predominance of passive symplasmic phloem loading and sugar transport with leaf ageing in Camellia oleifera. Horticulture Plant Journal, 9, 811–825.

Ye Z, Yu J, Yan W, Zhang J, Yang D, Yao G, Liu Z, Wu Y, Hou X. 2021. Integrative iTRAQ-based proteomic and transcriptomic analysis reveals the accumulation patterns of key metabolites associated with oil quality during seed ripening of Camellia oleifera. Horticulture Research, 8, 157.

Yue C, Cao H L, Wang L, Zhou Y H, Huang Y T, Hao X Y, Wang Y C, Wang B, Yang Y J, Wang X C. 2015. Effects of cold acclimation on sugar metabolism and sugar-related gene expression in tea plant during the winter season. Plant Molecular Biology, 88, 591–608

de Zelicourt A, Colcombet J, Hirt H. 2016. The role of MAPK modules and ABA during abiotic stress signaling. Trends in Plant Science, 21, 677–685.

Zeng Q L, Chen R F, Zhao X Q, Shen R F, Noguchi A, Shinmachi F, Hasegawa I. 2013. Aluminum could be transported via phloem in Camellia oleifera Abel. Tree Physiology, 33, 96–105.

Zhang C, Han L, Slewinski T L, Sun J, Zhang J, Wang Z Y, Turgeon R. 2014. Symplasmic phloem loading in poplar. Plant Physiology, 166, 306–313.

Zhang H, Cui X, Guo Y, Luo C, Zhang L. 2018. Picea wilsonii transcription factor NAC2 enhanced plant tolerance to abiotic stress and participated in RFCP1-regulated flowering time. Plant Molecular Biology, 98, 471–493.

Zhao Q, Ren Y R, Wang Q J, Yao Y X, You C X, Hao Y J. 2016. Overexpression of MdbHLH104 gene enhances the tolerance to iron deficiency in apple. Plant Biotechnology Journal, 14, 1633–1645.

Zheng M, Beck M, Müller J, Chen T, Wang X, Wang F, Wang Q, Wang Y, Baluška F, Logan D C. 2009. Actin turnover is required for myosin-dependent mitochondrial movements in Arabidopsis root hairs. PLoS ONE, 4, e5961.

Zhou J, Du B, Chen Y, Cao Y, Yu M, Zhang L. 2022. Integrative physiological and transcriptomic analysis reveals the transition mechanism of sugar phloem unloading route in Camellia oleifera fruit. International Journal of Molecular Sciences, 23, 4590.