Scientia Agricultura Sinica ›› 2026, Vol. 59 ›› Issue (14): 3147-3161.doi: 10.3864/j.issn.0578-1752.2026.14.012

• HORTICULTURE • Previous Articles     Next Articles

Prediction and Evaluation of Suitable Habitats for Carya illinoinensis in China Based on an Optimized MaxEnt Model and Land Use Types

ZHAO Ning1(), ZHANG JiYu2, WANG Rui3, YUAN ShuangJin3, CHEN JinHui4, LI ShaSha1, CHEN ZhiKun1()   

  1. 1 Xi'an Botanical Garden of Shaanxi Province (Institute of Botany of Shaanxi Province)/Shaanxi Key Laboratory of Qinling Ecological Security, Xi'an 710061
    2 Institute of Botany, Jiangsu Province and Chinese Academy of Sciences, Nanjing 210014
    3 Shaanxi Provincial Forestry Science and Technology Promotion and International Project Management Center, Xi'an 710082
    4 Ankang Hanbin District Forestry Bureau, Ankang 725000, Shaanxi
  • Received:2025-08-17 Accepted:2026-04-08 Online:2026-07-16 Published:2026-07-21
  • Contact: CHEN ZhiKun

Abstract:

【Objective】Carya illinoinensis (C. illinoinensis) (Pecan), as a characteristic agricultural and economic nut tree species, has been widely cultivated in multiple provinces across China in recent years. This study aimed to provide a scientific basis for the scientific layout of the C. illinoinensis industry by investigating and predicting its suitable habitats and land resources in China.【Method】First, based on the optimized Maximum Entropy (MaxEnt) model, 29 environmental variables, including climate, topography, and soil factors from 167 C. illinoinensis cultivation sites in China were analyzed. Then the dominant environmental factors were screened out, and the potential suitable habitats of C. illinoinensis in China under the current climate scenario were predicted. Second, the available cultivation area for C. illinoinensis was extracted and predicted by overlaying the forest distribution data of China.【Result】The MaxEnt model performed optimally with Feature Class (FC)=Linear-Quadratic (LQ) and Regularization Multiplier (RM) = 2. The AUC value of the ROC curve was 0.93, and the mean TSS of the optimized model was 0.7655, demonstrating that the optimized MaxEnt model exhibits exceptionally high reliability and accuracy. The suitable habitats of C. illinoinensis were mainly influenced by temperature and altitude. The primary dominant environmental factor was the minimum temperature of the coldest month (bio6), followed by altitude, isothermality (bio3), and temperature seasonality (bio4). The corresponding suitable threshold ranges were as follows: -5 ℃ to 5 ℃ for the minimum temperature of the coldest month (bio6), below 1 000 m for altitude, 15-30 and 50-57 for isothermality (bio3), and 400-1 000 for temperature seasonality (bio4). Ten provinces in China were identified as moderately to highly suitable habitats for C. illinoinensis, with a medium-suitable area of 9 208.69×104 hm2, accounting for 10% of China's total land area, and the high suitable area of 6 411.22×104 hm2, accounting for 7% of China's total land area. By overlaying with the forest distribution data of China, the actual available cultivation area for C. illinoinensis in China was 1 360.53×104 hm2.【Conclusion】In China, C. illinoinensis has potentially suitable habitats in Anhui, Hubei, Henan, Jiangsu, Hunan, Jiangxi, Zhejiang, the central part of Yunnan, south-central Shaanxi, and southwestern Shandong. After overlaying the suitable habitat map with China's forest land distribution data, the actual usable area was significantly reduced. At the provincial scale, Hubei Province had the largest area of highly suitable forest land, reaching 343.35×104 hm2, followed by Hunan Province with 305.57×104 hm2. Jiangxi and Anhui ranked third and fourth, with 219.81×104 and 161.83×104 hm2, respectively, making them important potential main producing areas for pecans. Combining the optimized MaxEnt model with land use types can effectively predict and scientifically evaluate the suitable areas for pecans in China, thereby providing a scientific basis for the development of the pecan industry in China.

Key words: Carya illinoinensis, MaxEnt model, suitable habitat prediction, environmental variables

Fig. 1

Sample points of the planting distribution of pecan in China"

Table 1

Environmental impact factors"

环境变量
Environment variable
代号
Code
环境因子名称
Name of environmental factor
单位
Unit
贡献率
Rate of contribution (%)
气候变量
Weather variable
Bio1 年均温 Mean annual temperature 4.7
Bio2 平均气温日较差 Mean diurnal temperature range 2.9
Bio3* 等温性(bio2 /bioc7)*100 Isothermal property ratio - 1.6
Bio4* 温度季节性*100 Standard deviation of seasonal variation of temperature - 0.8
Bio5 最暖月最高温 Max temperature of the warmest month 0.5
Bio6* 最冷月最低温 Minimum temperature of the coldest month 20.7
Bio7 气温年较差(bio5-bio6) Temperature annual range 0.4
Bio8 最湿季均温 Mean temperature of the wettest quarter 1.0
Bio9 最干季均温 Mean temperature of the direst quarter 0.9
Bio10 最热季均温 Mean temperature of the warmest quarter 0.7
Bio11 最冷季均温 Mean temperature of the coldest quarter 0.4
Bio12 年降水量 Annual precipitation mm 2.1
Bio13 最湿月降水量 Precipitation of the wettest month mm 0.3
Bio14* 最干月降水量 Precipitation of the driest month mm 43.7
Bio15* 降水季节性变动 Precipitation seasonality 1.5
Bio16 最湿季降水量 Precipitation of the wettest quarter mm 0.1
Bio17 最干季降水量 Precipitation of the driest quarter mm 2.2
Bio18 最暖季降水量 Precipitation of the warmest quarter mm 0.2
Bio19 最冷季降水量 Precipitation of the coldest quarter mm 0.5
地形变量
Landform variable
Dem* 海拔Altitude m 3.1
Slope* 坡度Slope 4.0
Aspect* 坡向Aspect 1.3
土壤变量
Soil variable
T_Clay* 顶层黏土含量Topsoil clay content % 1.3
T_Gravel* 顶层碎石体积百分比Topsoil gravel content (% by volume) % 1.2
T_PH* 顶层酸碱度(-log(H+))Topsoil pH(-log(H+)) 0.8
T_Sand* 顶层沙含量Topsoil sand content % 0.3
T_Silt* 顶层淤泥含量Topsoil silt content % 2.4
T_Oc* 顶层有机碳含量Topsoil organic carbon content % 0.2
T_Ref_Bulk 顶层土壤容重Topsoil bulk density kg·dm-3 0.1

Fig. 2

The correlation of environmental variables"

Table 2

The contribution rate and Permutation importance of climate variables"

环境因子
Climate variable
贡献率
Percent contribution
(%)
随机分布重要性
Permutation importance (%)
Bio6 82.0 74.0
Dem 7.1 10.5
T_Silt 2.9 0.5
Bio3 2.2 4.8
Bio15 1.6 2.2
Bio4 1.3 6.2
Slope 1.2 0.2
Aspect 0.4 0.1
T_Clay 0.4 0.2
T_PH 0.3 0.3
T_Gravel 0.3 0.3
T_Sand 0.3 0.7
Bio14 0 0
T_Oc 0 0

Fig. 3

Parameter selection and accuracy evaluation of the optimized MaxEnt model A:Selection of the MaxEnt model parameter; B: Results of the ROC precision test in the MaxEnt model; C: Average Omisslon and Predicted Area in the MaxEnt model In Figure A, H: Hinge; L: Linear; LQ: Linear+Quadratic; LQH: Linear+Quadratic+Hinge; LQHP: Linear+Quadratic+Hinge+Product; LQHPT: Linear+Quadratic+Hinge +Product+Threshold"

Fig. 4

Jackknife test to examine training gain result of climate variables"

Fig. 5

Response curves of major climatic factors"

Fig. 6

The division and scale of the suitable areas for pecans A: Division of suitable growing areas of pecan in China; B: Proportion statistics of suitable planting area of pecan; C: Suitable area of pecan in major provinces"

Fig. 7

The suitable growth classification and scale of pecan based on forest land resources A: Grading of suitable areas for pecans based on China’s forest land resources; B: The area of suitable growth zones for pecans in the forest areas of major provinces in China"

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