Ahmad A, Li W, Zhang H, Wang H, Wang P, Jiao Y, Zhao C, Yang G, Hong D. 2023. Linkage and association mapping of ovule number per ovary (ON) in oilseed rape (Brassica napus L.). Molecular Breeding, 43, 11.

Alonso-Blanco C, Blankestijn-de Vries H, Hanhart C J, Koornneef M. 1999. Natural allelic variation at seed size loci in relation to other life history traits of Arabidopsis thaliana. Proceedings of the National Academy of Sciences of the United States of America, 96, 4710–4717.

Armenta-Medina A, Gillmor C S. 2019. Genetic, molecular and parent-of-origin regulation of early embryogenesis in flowering plants. Current Topics in Developmental Biology, 131, 497–543.

Azhakanandam S, Nole-Wilson S, Bao F, Franks R G. 2008. SEUSS and AINTEGUMENTA mediate patterning and ovule initiation during gynoecium medial domain development. Plant Physiology, 146, 1165–1181.

Baker C C, Sieber P, Wellmer F, Meyerowitz E M. 2005. The early extra petals1 mutant uncovers a role for microRNA miR164c in regulating petal number in Arabidopsis. Current Biology, 15, 303–315.

Balasubramanian S, Schneitz K. 2000. NOZZLE regulates proximal–distal pattern formation, cell proliferation and early sporogenesis during ovule development in Arabidopsis thaliana. Development, 127, 4227–4238.

Balasubramanian S, Schneitz K. 2002. NOZZLE links proximal–distal and adaxial–abaxial pattern formation during ovule development in Arabidopsis thaliana. Development, 129, 4291–4300.

Banks H, Himanen I, Lewis G P. 2010. Evolution of pollen, stigmas and ovule numbers at the caesalpinioid-mimosoid interface (Fabaceae). Botanical Journal of the Linnean Society, 162, 594–615.

Bao F, Azhakanandam S, Franks R G. 2010. SEUSS and SEUSS-LIKE transcriptional adaptors regulate floral and embryonic development in Arabidopsis. Plant Physiology, 152, 821–836.

Barro-Trastoy D, Carrera E, Baños J, Palau-Rodríguez J, Ruiz-Rivero O, Tornero P, Alonso J M, López-Díaz I, Gómez M D, Perez-Amador M A. 2020a. Regulation of ovule initiation by gibberellins and brassinosteroids in tomato and Arabidopsis: Two plant species, two molecular mechanisms. The Plant Journal, 102, 1026–1041.

Barro-Trastoy D, Dolores Gómez M D, Tornero P, Perez-Amador M A. 2020b. On the way to ovules: The hormonal regulation of ovule development. Critical Reviews in Plant Sciences, 39, 431–456.

Bartrina I, Otto E, Strnad M, Werner T, Schmülling T. 2011. Cytokinin regulates the activity of reproductive meristems, flower organ size, ovule formation, and thus seed yield in Arabidopsis thaliana. The Plant Cell, 23, 69–80.

Basunanda P, Radoev M, Ecke W, Friedt W, Becker H, Snowdon R J. 2010. Comparative mapping of quantitative trait loci involved in heterosis for seedling and yield traits in oilseed rape (Brassica napus L.). Theoretical and Applied Genetics, 120, 271–281.

Becker A. 2020. A molecular update on the origin of the carpel. Current Opinion in Plant Biology, 53, 15–22.

Bencivenga S, Simonini S, Benková E, Colombo L. 2012. The transcription factors BEL1 and SPL are required for cytokinin and auxin signaling during ovule development in Arabidopsis. The Plant Cell, 24, 2886–2897.

Benková E, Michniewicz M, Sauer M, Teichmann T, Seifertová D, Jürgens G, Friml J. 2003. Local, efflux-dependent auxin gradients as a common module for plant organ formation. Cell, 115, 591–602.

van Berkel K, de Boer R J, Scheres B, ten Tusscher K. 2013. Polar auxin transport: Models and mechanisms. Development, 140, 2253–2268.

Bilsborough G D, Runions A, Barkoulas M, Jenkins H W, Hasson A, Galinha C, Laufs P, Hay A, Prusinkiewicz P, Tsiantis M. 2011. Model for the regulation of Arabidopsis thaliana leaf margin development. Proceedings of the National Academy of Sciences of the United States of America, 108, 3424–3429.

Brambilla V, Battaglia R, Colombo M, Masiero S, Bencivenga S, Kater M M, Colombo L. 2007. Genetic and molecular interactions between BELL1 and MADS box factors support ovule development in Arabidopsis. The Plant Cell, 19, 2544–2556.

Brumos J, Robles L M, Yun J, Vu T C, Jackson S, Alonso J M, Stepanova A N. 2018. Local auxin biosynthesis is a key regulator of plant development. Developmental Cell, 47, 306–318.

Ceccato L, Masiero S, Sinha Roy D, Bencivenga S, Roig-Villanova I, Ditengou F A, Palme K, Simon R, Colombo L. 2013. Maternal control of PIN1 is required for female gametophyte development in Arabidopsis. PLoS ONE, 8, e66148.

Cheng Y, Dai X, Zhao Y. 2006. Auxin biosynthesis by the YUCCA flavin monooxygenases controls the formation of floral organs and vascular tissues in Arabidopsis. Genes & Development, 20, 1790–1799.

Cucinotta M, Colombo L, Roig-Villanova I J. 2014. Ovule development, a new model for lateral organ formation. Frontiers in Plant Science, 5, 117.

Cucinotta M, Di Marzo M, Guazzotti A, de Folter S, Kater M M, Colombo L. 2020. Gynoecium size and ovule number are interconnected traits that impact seed yield. Journal of Experimental Botany, 71, 2479–2489.

Cucinotta M, Manrique S, Cuesta C, Benkova E, Novak O, Colombo L. 2018. CUP-SHAPED COTYLEDON1 (CUC1) and CUC2 regulate cytokinin homeostasis to determine ovule number in Arabidopsis. Journal of Experimental Botany, 69, 5169–5176.

Cucinotta M, Manrique S, Guazzotti A, Quadrelli N E, Mendes M A, Benkova E, Colombo L. 2016. Cytokinin response factors integrate auxin and cytokinin pathways for female reproductive organ development. Development, 143, 4419–4424.

Davière J M, Achard P. 2016. A pivotal role of DELLAs in regulating multiple hormone signals. Molecular Plant, 9, 10–20.

Enugutti B, Kirchhelle C, Oelschner M, Torres Ruiz R A, Schliebner I, Leister D, Schneitz K. 2012. Regulation of planar growth by the Arabidopsis AGC protein kinase UNICORN. Proceedings of the National Academy of Sciences of the United States of America, 109, 15060–15065.

Enugutti B, Schneitz K. 2013. Genetic analysis of ectopic growth suppression during planar growth of integuments mediated by the Arabidopsis AGC protein kinase UNICORN. BMC Plant Biology, 13, 2.

Fridman Y, Savaldi-Goldstein S. 2013. Brassinosteroids in growth control: How, when and where. Plant Science, 209, 24–31.

Galbiati F, Sinha Roy D, Simonini S, Cucinotta M, Ceccato L, Cuesta C, Simaskova M, Benkova E, Kamiuchi Y, Aida M, Weijers D. 2013. An integrative model of the control of ovule primordia formation. The Plant Journal, 76, 446–455.

Gallego-Giraldo C, Hu J, Urbez C, Gómez M D, Sun T P, Perez-Amador M A. 2014. Role of the gibberellin receptors GID 1 during fruit-set in Arabidopsis. The Plant Journal, 79, 1020–1032.

Gómez M D, Barro-Trastoy D, Escoms E, Saura-Sánchez M, Sánchez I, Briones-Moreno A, Vera-Sirera F, Carrera E, Ripoll J J, Yanofsky M F, Lopez-Diaz I, Alonso J M, Perez-Amador M A. 2018. Gibberellins negatively modulate ovule number in plants. Development, 145, dev163865.

Gómez M D, Barro-Trastoy D, Fuster-Almunia C, Tornero P, Alonso J M, Perez-Amador M A. 2020. Gibberellin-mediated RGA-LIKE1 degradation regulates embryo sac development in Arabidopsis. Journal of Experimental Botany, 71, 7059–7072.

Gómez M D, Fuster-Almunia C, Ocana-Cuesta J, Alonso J M, Perez-Amador M A. 2019. RGL2 controls flower development, ovule number and fertility in Arabidopsis. Plant Science, 281, 82–92.

Gómez M D, Urbez C, Perez-Amador M A, Carbonell J. 2011. Characterization of constricted fruit (ctf) mutant uncovers a role for AtMYB117/LOF1 in ovule and fruit development in Arabidopsis thaliana. PLoS ONE, 6, e18760.

Gómez M D, Ventimilla D, Sacristan R, Perez-Amador M A. 2016. Gibberellins regulate ovule integument development by interfering with the transcription factor ATS. Plant Physiology, 172, 2403–2415.

Gonçalves B, Hasson A, Belcram K, Cortizo M, Morin H, Nikovics K, Vialette-Guiraud A, Takeda S, Aida M, Laufs P, Arnaud N. 2015. A conserved role for CUP-SHAPED COTYLEDON genes during ovule development. The Plant Journal, 83, 732–742.

Hashida T, Nakatsuji R, Budahn H, Schrader O, Peterka H, Fujimura T, Kubo N, Hirai M. 2013. Construction of a chromosome-assigned, sequence-tagged linkage map for the radish, Raphanus sativus L. and QTL analysis of morphological traits. Breeding Science, 63, 218–226.

Hashimoto K, Miyashima S, Sato-Nara K, Yamada T, Nakajima K. 2018. Functionally diversified members of the MIR165/6 gene family regulate ovule morphogenesis in Arabidopsis thaliana. Plant and Cell Physiology, 59, 1017–1026.

Heisler M G, Byrne M E. 2020. Progress in understanding the role of auxin in lateral organ development in plants. Current Opinion in Plant Biology, 53, 73–79.

Huang H Y, Jiang W B, Hu Y W, Wu P, Zhu J Y, Liang W Q, Wang Z Y, Lin W H. 2013. BR signal influences Arabidopsis ovule and seed number through regulating related genes expression by BZR1. Molecular Plant, 6, 456–469.

Hwang I, Sheen J, Müller B. 2012. Cytokinin signaling networks. Annual Review of Plant Biology, 63, 353–380.

Ishida T, Aida M, Takada S, Tasaka M. 2000. Involvement of CUP-SHAPED COTYLEDON genes in gynoecium and ovule development in Arabidopsis thaliana. Plant and Cell Physiology, 41, 60–67.

Jia D, Chen L G, Yin G, Yang X, Gao Z, Guo Y, Sun Y, Tang W. 2020. Brassinosteroids regulate outer ovule integument growth in part via the control of INNER NO OUTER by BRASSINOZOLE-RESISTANT family transcription factors. Journal of Integrative Plant Biology, 62, 1093–1111.

Jiao Y, Zhang K, Cai G, Yu K, Amoo O, Han S, Zhao X, Zhang H, Hu L, Wang B, Fan C, Zhou Y. 2021. Fine mapping and candidate gene analysis of a major locus controlling ovule abortion and seed number per silique in Brassica napus L. Theoretical and Applied Genetics, 134, 2517–2530.

Jung J H, Park C M. 2007. MIR166/165 genes exhibit dynamic expression patterns in regulating shoot apical meristem and floral development in Arabidopsis. Planta, 225, 1327–1338.

Kawamoto N, Del Carpio D P, Hofmann A, Mizuta Y, Kurihara D, Higashiyama T, Uchida N, Torii K U, Colombo L, Groth G, Simon R. 2020. A peptide pair coordinates regular ovule initiation patterns with seed number and fruit size. Current Biology, 30, 4352–4361.

Kelley D R, Arreola A, Gallagher T L, Gasser C S. 2012. ETTIN (ARF3) physically interacts with KANADI proteins to form a functional complex essential for integument development and polarity determination in Arabidopsis. Development, 139, 1105–1109.

Kelley D R, Gasser C S. 2009. Ovule development: Genetic trends and evolutionary considerations. Sexual Plant Reproduction, 22, 229–234.

Khan S U, Yangmiao J, Liu S, Zhang K, Khan M H U, Zhai Y, Olalekan A, Fan C, Zhou Y. 2019. Genome-wide association studies in the genetic dissection of ovule number, seed number, and seed weight in Brassica napus L. Industrial Crops and Products, 142, 111877.

Klucher K M, Chow H, Reiser L, Fischer R L.1996. The AINTEGUMENTA gene of Arabidopsis required for ovule and female gametophyte development is related to the floral homeotic gene APETALA2. The Plant Cell, 8, 137–153.

Kozlowski J, Stearns S C. 1989. Hypotheses for the production of excess zygotes: Models of bet-hedging and selective abortion. Evolution, 43, 1369–1377.

Krizek B A, Blakley I C, Ho Y Y, Freese N, Loraine A E. 2020. The Arabidopsis transcription factor AINTEGUMENTA orchestrates patterning genes and auxin signaling in the establishment of floral growth and form. The Plant Journal, 103, 752–768.

Laufs P, Peaucelle A, Morin H, Traas J. 2004. MicroRNA regulation of the CUC genes is required for boundary size control in Arabidopsis meristems. Development, 131, 4311–4322.

Leyser O. 2018. Auxin signaling. Plant Physiology, 176, 465–479.

Li B F, Yu S X, Hu L Q, Zhang Y J, Zhai N, Xu L, Lin W H. 2018. Simple culture methods and treatment to study hormonal regulation of ovule development. Frontiers in Plant Science, 9, 784.

Li L C, Qin G J, Tsuge T, Hou X H, Ding M Y, Aoyama T, Oka A, Chen Z, Gu H, Zhao Y, Qu L J. 2008. SPOROCYTELESS modulates YUCCA expression to regulate the development of lateral organs in Arabidopsis. New Phytologist, 179, 751–764.

Liao S, Wang L, Li J, Ruan Y L. 2020. Cell wall invertase is essential for ovule development through sugar signaling rather than provision of carbon nutrients. Plant Physiology, 183, 1126–1144.

Liu Z, Franks R G, Klink V P. 2000. Regulation of gynoecium marginal tissue formation by LEUNIG and AINTEGUMENTA. The Plant Cell, 12, 1879–1891.

Lohmann D, Stacey N, Breuninger H, Jikumaru Y, Müller D, Sicard A, Leyser O, Yamaguchi S, Lenhard M. 2010. SLOW MOTION is required for within-plant auxin homeostasis and normal timing of lateral organ initiation at the shoot meristem in Arabidopsis. The Plant Cell, 22, 335–348.

Mallory A C, Dugas D V, Bartel D P, Bartel B. 2004. MicroRNA regulation of NAC-domain targets is required for proper formation and separation of adjacent embryonic, vegetative, and floral organs. Current Biology, 14, 1035–1046.

Marsch-Martínez N, de Folter S. 2016. Hormonal control of the development of the gynoecium. Current Opinion in Plant Biology, 29, 104–114.

Marsch-Martínez N, Ramos-Cruz D, Irepan Reyes-Olalde J, Lozano-Sotomayor P, Zúñiga-Mayo V M, de Folter S. 2012. The role of cytokinin during Arabidopsis gynoecia and fruit morphogenesis and patterning. The Plant Journal, 72, 222–234.

Matilla A J. 2019. Seed coat formation: Its evolution and regulation. Seed Science Research, 29, 215–226.

McAbee J M, Hill T A, Skinner D J, Izhaki A, Hauser B A, Meister R J, Venugopala Reddy G, Meyerowitz E M, Bowman J L, Gasser C S. 2006. ABERRANT TESTA SHAPE encodes a KANADI family member, linking polarity determination to separation and growth of Arabidopsis ovule integuments. The Plant Journal, 46, 522–531.

Meister R J, Kotow L M, Gasser C S. 2002. SUPERMAN attenuates positive INNER NO OUTER autoregulation to maintain polar development of Arabidopsis ovule outer integuments. Development, 129, 4281–4289.

Mizukami Y, Fischer R L. 2000. Plant organ size control: AINTEGUMENTA regulates growth and cell numbers during organogenesis. Proceedings of the National Academy of Sciences of the United States of America, 97, 942–947.

Morffy N, Strader L C. 2020. Old Town Roads: Routes of auxin biosynthesis across kingdoms. Current Opinion in Plant Biology, 55, 21–27.

Mutte S K, Kato H, Rothfels C, Melkonian M, Wong G K, Weijers D. 2018. Origin and evolution of the nuclear auxin response system. eLife, 7, e33399.

Nahar M A U, Ishida T, Smyth D R, Tasaka M, Aida M. 2012. Interactions of CUP-SHAPED COTYLEDON and SPATULA genes control carpel margin development in Arabidopsis thaliana. Plant and Cell Physiology, 53, 1134–1143.

Nemhauser J L, Feldman L J, Zambryski P C. 2000. Auxin and ETTIN in Arabidopsis gynoecium morphogenesis. Development, 127, 3877–3888.

Nole-Wilson S, Azhakanandam S, Franks R G. 2010. Polar auxin transport together with AINTEGUMENTA and REVOLUTA coordinate early Arabidopsis gynoecium development. Developmental Biology, 346, 181–195.

Petrella R, Gabrieli F, Cavalleri A, Schneitz K, Colombo L, Cucinotta M J D. 2022. Pivotal role of STIP in ovule pattern formation and female germline development in Arabidopsis thaliana. Development, 149, dev201184.

Phillips A R, Evans M M. 2020. Maternal regulation of seed growth and patterning in flowering plants. Current Topics in Developmental Biology, 140, 257–282.

Pinyopich A, Ditta, G S, Savidge B, Liljegren S J, Baumann E, Wisman E, Yanofsky M F. 2003. Assessing the redundancy of MADS-box genes during carpel and ovule development. Nature, 424, 85–88.

Planas-Riverola A, Gupta A, Betegón-Putze I, Bosch N, Ibañes M, Caño-Delgado A I. 2019. Brassinosteroid signaling in plant development and adaptation to stress. Development, 146, dev151894.

Qadir M, Qin L, Ye J, Ahmad N, Wang X, Shi J, Wang H. 2022. Genetic dissection of the natural variation of ovule number per ovary in oilseed rape germplasm (Brassica napus L.). Frontiers in Plant Science, 13, 999790.

Qadir M, Wang X, Shah S R U, Zhou X R, Shi J, Wang H. 2021. Molecular network for regulation of ovule number in plants. International Journal of Molecular Sciences, 22, 12965.

Raboanatahiry N, Chao H, He J, Li H, Yin Y, Li M. 2022. Construction of a quantitative genomic map, identification and expression analysis of candidate genes for agronomic and disease-related traits in Brassica napus. Frontiers in Plant Science, 13, 862363.

Reiser L, Modrusan Z, Margossian L, Samach A, Ohad N, Haughn G W, Fischer R L. 1995. The BELL1 gene encodes a homeodomain protein involved in pattern formation in the Arabidopsis ovule primordium. Cell, 83, 735–742.

Reyes-Olalde J I, Zúñiga-Mayo V M, Serwatowska J, Chavez Montes R A, Lozano-Sotomayor P, Herrera-Ubaldo H, Gonzalez-Aguilera K L, Ballester P, Ripoll J J, Ezquer I, Paolo D, Heyl A, Colombo L, Yanofsky M F, Ferrandiz C, Marsch-Martínez N, de Folter S. 2017. The bHLH transcription factor SPATULA enables cytokinin signaling, and both activate auxin biosynthesis and transport genes at the medial domain of the gynoecium. PLoS Genetics, 13, e1006726.

Rhoades M W, Reinhart B J, Lim L P, Burge C B, Bartel B, Bartel D P. 2002. Prediction of plant microRNA targets. Cell, 110, 513–520.

Rizza A, Jones A M. 2019. The makings of a gradient: Spatiotemporal distribution of gibberellins in plant development. Current Opinion in Plant Biology, 47, 9–15.

Rodríguez-Cazorla E, Ortuño-Miquel S, Candela H, Bailey-Steinitz L J, Yanofsky M F, Martínez-Laborda A, Ripoll J J, Vera A. 2018. Ovule identity mediated by pre-mRNA processing in Arabidopsis. PLoS Genetics, 14, e1007182.

Rodríguez-Cazorla E, Ripoll J J, Ortuño-Miquel S, Martínez-Laborda A, Vera A. 2020. Dissection of the Arabidopsis HUA-PEP gene activity reveals that ovule fate specification requires restriction of the floral A-function. New Phytologist, 227, 1222–1234.

Schneitz K, Balasubramanian S, Schiefthaler U. 1998. Organogenesis in plants: The molecular and genetic control of ovule development. Trends in Plant Science, 3, 468–472.

Schneitz K, Hülskamp M, Pruitt R E. 1995. Wild-type ovule development in Arabidopsis thaliana: A light microscope study of cleared whole-mount tissue. The Plant Journal, 7, 731–749.

Shi J, Li R, Qiu D, Jiang C, Long Y, Morgan C, Bancroft I, Zhao J, Meng J. 2009. Unraveling the complex trait of crop yield with quantitative trait loci mapping in Brassica napus. Genetics, 182, 851–861.

Shirley N J, Aubert M K, Wilkinson L G, Bird D C, Lora J, Yang X, Tucker M R. 2019. Translating auxin responses into ovules, seeds and yield: Insight from Arabidopsis and the cereals. Plant Biology, 61, 310–336.

Šimášková M, O’Brien J A, Khan M, Van Noorden G, Ötvös K, Vieten A, De Clercq I, Van Haperen J M A, Cuesta C, Hoyerová K, Vanneste S, Marhavý P, Wabnik K, Van Breusegem F, Nowack M, Murphy A, Friml J, Weijers D, Beeckman T, Benková E. 2015. Cytokinin response factors regulate PIN-FORMED auxin transporters. Nature Communications, 6, 8717.

Simon M K, Skinner D J, Gallagher T L, Gasser C S. 2017. Integument development in Arabidopsis depends on interaction of YABBY protein INNER NO OUTER with coactivators and corepressors. Genetics, 207, 1489–1500.

Smyth D R, Bowman J L, Meyerowitz E M. 1990. Early flower development in Arabidopsis. The Plant Cell, 2, 755–767.

Sohlberg J J, Myrenås M, Kuusk S, Lagercrantz U, Kowalczyk M, Sandberg G, Sundberg E. 2006. STY1 regulates auxin homeostasis and affects apical-basal patterning of the Arabidopsis gynoecium. The Plant Journal, 47, 112–123.

Tischner T, Allphin L, Chase K, Orf J H, Lark K G. 2003. Genetics of seed abortion and reproductive traits in soybean. Crop Science, 43, 464–473.

Villanueva J M, Broadhvest J, Hauser B A, Meister R J, Schneitz K, Gasser C S. 1999. INNER NO OUTER regulates abaxial–adaxial patterning in Arabidopsis ovules. Genes & Development, 13, 3160–3169.

Wang Y, Jiao Y. 2018. Auxin and above-ground meristems. Journal of Experimental Botany, 69, 147–154.

Weijers D, Nemhauser J, Yang Z. 2018. Auxin: Small molecule, big impact. Journal of Experimental Botany, 69, 133–136.

Wynn A N, Seaman A A, Jones A L, Franks R G. 2014. Novel functional roles for PERIANTHIA and SEUSS during floral organ identity specification, floral meristem termination, and gynoecial development. Frontiers in Plant Science, 5, 130.

Yamada T, Sasaki Y, Hashimoto K, Nakajima K, Gasser C S. 2016. CORONA, PHABULOSA and PHAVOLUTA collaborate with BELL1 to confine WUSCHEL expression to the nuceLlus in Arabidopsis ovules. Development, 143, 422–426.

Yamaguchi N, Wu M F, Winter C M, Berns M C, Nole-Wilson S, Yamaguchi A, Coupland G, Krizek B A, Wagner D. 2013. A molecular framework for auxin-mediated initiation of flower primordia. Developmental Cell, 24, 271–282.

Yu S X, Jiang Y T, Lin W H. 2022. Ovule initiation: the essential step controlling offspring number in Arabidopsis. Journal of Integrated Plant Biology, 64, 1469–1486.

Yuan J, Kessler S A. 2019. A genome-wide association study reveals a novel regulator of ovule number and fertility in Arabidopsis thaliana. PLoS Genetics, 15, e1007934.

Žádníková P, Simon R. 2014. How boundaries control plant development. Current Opinion in Plant Biology, 17, 116–125.

Zhang L, Yang G, Liu P, Hong D, Li S, He Q. 2011. Genetic and correlation analysis of silique-traits in Brassica napus L. by quantitative trait locus mapping. Theoretical and Applied Genetics, 122, 21–31.

Zhou J J, Luo J. 2018. The PIN-FORMED auxin efflux carriers in plants. International Journal of Molecular Sciences, 19, 2759.

Zu S H, Jiang Y T, Chang J H, Zhang Y J, Xue H W, Lin W H. 2022. Interaction of brassinosteroid and cytokinin promotes ovule initiation and increases seed number per silique in Arabidopsis. Journal of Integrative Plant Biology, 64, 702–716.