Decreased panicle N application alleviates the negative effects of
shading on rice grain yield and grain quality

doi:10.1016/j.jia.2022.08.045

Journal of Integrative Agriculture

2023, Vol. 22

Issue (7): 2041-2053 DOI: 10.1016/j.jia.2022.08.045

Crop Science

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Decreased panicle N application alleviates the negative effects of shading on rice grain yield and grain quality

WEI Huan-he¹, GE Jia-lin¹, ZHANG Xu-bin¹, ZHU Wang^1, ², DENG Fei³, REN Wan-jun³, CHEN Ying-long¹, MENG Tian-yao^2#, DAI Qi-gen^1, ^2#

¹Jiangsu Key Laboratory of Crop Cultivation and Physiology/Jiangsu Key Laboratory of Crop Genetics and Physiology/Jiangsu Co-Innovation Center for Modern Production Technology of Grain Crops/Research Institute of Rice Industrial Engineering Technology/College of Agriculture, Yangzhou University, Yangzhou 225009, P.R.China
² Institutes of Agricultural Sciences and Technology Development/Joint International Research Laboratory of Agriculture and Agri-Product Safety, Ministry of Education/College of Agriculture, Yangzhou University, Yangzhou 225009, P.R.China
³ Crop Ecophysiology and Cultivation Key Laboratory of Sichuan Province/Key Laboratory of Crop Ecophysiology and Farming System in Southwest China, Ministry of Agriculture and Rural Affairs/College of Agronomy, Sichuan Agricultural University, Chengdu 611130, P.R.China

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摘要

弱光胁迫已成为水稻生产中日趋严重的逆境，然而关于弱光胁迫对华东地区水稻产量和品质的研究报道仍较少；此外，也应探索合理的氮肥施用措施以减轻弱光胁迫对水稻产量和品质的负面效应。本研究旨在明确遮光（无遮光、花后遮光）和穗肥施用量（N_DP，降低穗肥用量；N_M_P，正常穗肥用量；N_I_P，增加穗肥用量）对水稻产量和品质相关性状的影响。与无遮光相比，遮光处理下水稻减产9.5%-14.8% (P<0.05)，主要是由于较低的结实率和粒重。无遮光处理下，N_MP和N_IP的产量显著高于(P<0.05)N_DP；遮光处理下，不同穗肥施用处理的产量则无显著差异。与N_MP和N_IP相比，N_DP在遮光处理的减产幅度小，这主要是由于其较高的结实率和粒重。与无遮光相比，遮光处理降低了花后叶片光合速率、成熟期干物质、抽穗至成熟期干物质积累量和成熟期茎鞘非结构性碳水化合物(NSC)的积累量。N_DP则提高了遮光处理下的收获指数和NSC运转效率。与无遮光相比，遮光处理降低了稻米糙米率、精米率、整精米率和直链淀粉含量，提高了垩白米率、垩白面积和垩白度以及蛋白质含量。无遮光处理下稻米加工品质以N_MP最优，遮光处理下则以N_D_P最优。在无遮光和遮光处理下，N_D_P的垩白米率、垩白面积和垩白度均低于N_MP和N_IP。遮光处理下，N_D_P的直链淀粉含量和崩解值降低，蛋白质含量和消减值则提高，从而实现了与无遮光处理相近的食味值。本研究结果表明，花后遮光显著恶化水稻产量和稻米品质。遮光处理下，N_D_P可促进NSC转运、提高收获指数以及库容充实效率，从而降低产量损失；此外，N_D_P可维持遮光处理下稻米加工、外观和蒸煮食味品质。因此，适当降低穗肥施用可减轻弱光胁迫对水稻产量和品质的负面效应。

Abstract Light deficiency is a growing abiotic stress in rice production. However, few studies focus on shading effects on grain yield and quality of rice in East China. It is also essential to investigate proper nitrogen (N) application strategies that can effectively alleviate the negative impacts of light deficiency on grain yield and quality in rice. A two-year field experiment was conducted to explore the effects of shading (non-shading and shading from heading to maturity) and panicle N application (N_DP, decreased panicle N rate; N_MP, medium panicle N rate; N_IP, increased panicle N rate) treatments on rice yield- and quality-related characteristics. Compared with non-shading, shading resulted in a 9.5–14.8% yield loss (P<0.05), mainly due to lower filled-grain percentage and grain weight. N_MP and N_IP had higher (P<0.05) grain yield than N_DP under non-shading, and no significant difference was observed in rice grain yield among N_DP, N_MP, and N_IP under shading. Compared with N_MP and N_IP, N_DP achieved less yield loss under shading because of the increased filled-grain percentage and grain weight. Shading reduced leaf photosynthetic rate after heading, as well as shoot biomass weight at maturity, shoot biomass accumulation from heading to maturity, and nonstructural carbohydrate (NSC) content in the stem at maturity (P<0.05). The harvest index and NSC remobilization reserve of N_DP were increased under shading. Shading decreased (P<0.05) percentages of brown rice, milled rice, head rice, and amylose content while increasing (P<0.05) chalky rice percentage, chalky area, chalky degree, and grain protein. N_MP demonstrated a better milling quality under non-shading, while N_DP demonstrated under shading. N_DP exhibited both lower chalky rice percentage, chalky area, and chalky degree under non-shading and shading, compared with N_MP and N_IP. N_DP under shading decreased amylose content and breakdown but increased grain protein content and setback, contributing to similar overall palatability to nonshading. Our results suggested severe grain yield and quality penalty of rice when subjected to shading after heading. N_DP improved NSC remobilization, harvest index, and sink-filling efficiency and alleviated yield loss under shading. Besides, N_DP would maintain rice’s milling, appearance, and cooking and eating qualities under shading. Proper N management with a decreased panicle N rate could be adopted to mitigate the negative effects of shading on rice grain yield and quality.

Keywords: rice grain yield grain quality shading panicle N application

Received: 05 May 2022 Accepted: 24 June 2022

Fund:

This work was financed by the Joints Funds of the National Natural Science Foundation of China (U20A2022), the National Natural Science Foundation of China (31901448 and 32001466), the Postdoctoral Research Foundation of China (2020M671628 and 2020M671629), the Natural Science Foundation of the Jiangsu Higher Education Institutions, China (19KJB210004), the Key Research and Development Program of Jiangsu Province (BE2019343), and the Priority Academic Program Development of Jiangsu Higher Education Institutions.

About author: WEI Huan-he, E-mail: 006931@yzu.edu.cn; #Correspondence MENG Tian-yao, E-mail: 516060030@qq.com; DAI Qi-gen, E-mail: qgdai@yzu.edu.cn

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WEI Huan-he, GE Jia-lin, ZHANG Xu-bin, ZHU Wang, DENG Fei, REN Wan-jun, CHEN Ying-long, MENG Tian-yao, DAI Qi-gen. 2023. Decreased panicle N application alleviates the negative effects of shading on rice grain yield and grain quality. Journal of Integrative Agriculture, 22(7): 2041-2053.

Akhtar K, Wang W Y, Khan A, Ren G X, Afridi M Z, Feng Y Z, Yang G. 2019. Wheat straw mulching offset soil moisture deficient for improving physiological and growth performance of summer sown soybean. Agricultural Water Management, 211, 16–25.

Ambardekar A A, Siebenmorgen T J, Counce P A, Lanning S B, Mauromoustakos A. 2011. Impact of field-scale nighttime air temperatures during kernel development on rice milling quality. Field Crops Research, 122, 179–185.

Bahuguna R N, Solis C A, Shi W J, Jagadish K S V. 2017. Post-flowering night respiration and altered sink activity account for high night temperature-induced grain yield and quality loss in rice (Oryza sativa L.). Physiologia Plantarum, 159, 59–73.

Balindong J L, Ward R M, Liu L, Rose T J, Pallas L A, Ovenden B W, Snell P J, Waters D L E. 2018. Rice grain protein composition influences instrumental measures of rice cooking and eating quality. Journal of Cereal Science, 79, 35–42.

Bian J L, Xu F F, Han C, Qiu S, Ge J L, Xu J, Zhang H C, Wei H Y. 2018. Effects of planting methods on yield and quality of different types of japonica rice in northern Jiangsu plain, China. Journal of Integrative Agriculture, 17, 2624–2635.

Carrijo D R, Akbar N, Reis A F B, Li C Y, Gaudin A C M, Parikh S J, Green P G, Linquist B A. 2018. Impacts of variable soil drying in alternate wetting and drying rice systems on yields, grain arsenic concentration and soil moisture dynamics. Field Crops Research, 222, 101–110.

Chen H, Li Q P, Zeng Y L, Deng F, Ren W J. 2019. Effect of different shading materials on grain yield and quality of rice. Scientific Reports, 9, 9992.

Deng F, Li Q P, Chen H, Zeng Y L, Li B, Zhong X Y, Wang L, Ren W J. 2021a. Relationship between chalkiness and the structural and thermal properties of rice starch after shading during grain-filling stage. Carbohydrate Polymers, 252, 117212.

Deng F, Wang L, Pu S L, Mei X F, Li S X, Li Q P, Ren W J. 2018. Shading stress increases chalkiness by postponing caryopsis development and disturbing starch characteristics of rice grains. Agricultural and Forest Meteorology, 263, 49–58.

Deng F, Zeng Y L, Li Q P, He C Y, Li B, Zhu Y Y, Zhou X, Yang F, Zhong X Y, Wang L, Chen H, Zhou W, Ren W J. 2021b. Decreased anther dehiscence contributes to a lower fertilization rate of rice subjected to shading stress. Field Crops Research, 273, 108291.

Fabre D, Adriani D E, Dingkuhn M, Ishimaru T, Punzalan B, Lafarge T, Clement-Vidal A, Luquet D. 2016. The qTSN4 effect on flag leaf size, photosynthesis and panicle size, benefits to plant grain production in rice, depending on light availability. Frontiers in Plant Science, 7, 623.

Fang X M, Li Y S, Nie J, Wang C, Huang K H, Zhang Y K, Zhang Y L, She H Z, Liu X B, Ruan R W, Yuan X H, Yi Z L. 2018. Effects of nitrogen fertilizer and planting density on the leaf photosynthetic characteristics, agronomic traits and grain yield in common buckwheat (Fagopyrum esculentum M.). Field Crops Research, 219, 160–168.

Fitzgerald M Z, McCouch S R, Hall R D. 2009. Not just a grain of rice: the quest for quality. Trends in Plant Science, 14, 133–139.

Fu P H, Wang J, Zhang T, Huang J L, Peng S B. 2019. High nitrogen input causes poor grain filling of spikelets at the panicle base of super hybrid rice. Field Crops Research, 244, 107635.

GB/T 17891-2017. 2017. High Quality Paddy. National Standard of the People’s Republic of China. pp. 1–10. (in Chinese)

Grigg B C, Siebenmorgen T J, Norman R J. 2016. Effects of nitrogen rate and harvest moisture content on physicochemical properties and milling yields of rice. Cereal Chemistry, 93, 172–181.

He Y Y, Wang K C, Zhou C L, Wild M. 2018. A revisit of global dimming and brightening based on the sunshine duration. Geophysical Research Letters, 45, 4281–4289.

Hori K, Suzuki K, Lijima K, Ebana K. 2016. Variation in cooking and eating quality traits in Japanese rice germplasm accessions. Breeding Science, 66, 309–318.

Idowu O, Wang Y Z, Homma K, Nakazaki T, Xu Z J, Shiraiwa T. 2022. Interaction of erect panicle genotype and nitrogen fertilizer application on the source–sink ratio and nitrogen use efficiency in rice. Field Crops Research, 278, 108430.

Impa S M, Raju B, Hein N T, Sandhu J, Prasad P V V, Walia H, Jagadish S V K. 2021. High night temperature effects on wheat and rice: Current status and way forward. Plant, Cell & Environment, 44, 2049–2065.

Ishibashi Y, Okamura K, Miyazaki M, Phan T, Yuasa T, Iwaya-Inoue M. 2014. Expression of rice sucrose transporter gene OsSUT1 in sink and source organs shaded during grain filling may affect grain yield and quality. Environmental and Experimental Botany, 97, 49–54.

Jiang Q, Du Y L, Tian X Y, Wang Q S, Xiong R H, Xu G C, Yan C, Ding Y F. 2016. Effect of panicle nitrogen on grain filling characteristics of high-yielding rice cultivars. European Journal of Agronomy, 74, 185–192.

Kakar K, Nitta Y, Asagi N, Komatsuzaki M, Shiotsu F, Kokubo T, Xuan T D. 2019. Morphological analysis on comparison of organic and chemical fertilizers on grain quality of rice at different planting densities. Plant Production Science, 22, 510–518.

Kamiji Y, Yoshida H, Palta J A, Sakuratani T, Shiraiwa T. 2011. N applications that increase plant N during panicle development are highly effective in increasing spikelet number in rice. Field Crops Research, 122, 242–247.

Khazaei H, Wach D, Pecio A, Vandenberg A, Stoddard F L. 2019. Genetic analysis of photosynthesis-related traits in faba bean (Vicia faba) for crop improvement. Plant Breeding, 138, 761–769.

Kobata T, Sugawara M, Takatu S. 2000. Shading during the early grain filling period does not affect potential grain dry matter increase in rice. Agronomy Journal, 92, 411–417.

Li H Y, Yan S, Yang L, Xu M H, Ji J Y, Mao H J, Song Y J, Wang J, Sun B G. 2021. Starch gelatinization in the surface layer of rice grains is crucial in reducing the stickiness of parboiled rice. Food Chemistry, 341, 128202.

Li Q P, Deng F, Chen H, Zeng Y L, Li B, Zhong X Y, Wang L, Zhou W, Chen Y, Ren W J. 2020. Shading decreases rice yield by impeding grain-filling progress after heading. Agronomy Journal, 112, 4018–4030.

Liu K, Yang R, Lu J, Wang X Y, Lu B L, Tian X H, Zhang Y B. 2019. Radiation use efficiency and source–sink changes of super hybrid rice under shade stress during grain-filling stage. Agronomy Journal, 111, 1788–1798.

Mishra J S, Poonia S P, Kumar R, Dubey R, Kumar V, Mondal S, Dwivedi S K, Rao K K, Kumar R, Tamta M, Verma M, Saurabh K, Kumar S, Bhatt B P, Malik R K, Mcdonald A, Bhaskar S. 2021. An impact of agronomic practices of sustainable rice–wheat crop intensification on food security, economic adaptability, and environmental mitigation across eastern Indo-Gangetic Plains. Field Crops Research, 267, 108164.

Mo Z W, Li W, Pan S G, Fitzgerald T L, Xiao F, Tang Y J, Wang Y L, Duan M Y, Tian H, Tang X R. 2015. Shading during the grain filling period increases 2-acetyl-1-pyrroline content in fragrant rice. Rice, 8, 9.

Morita S, Nakano H. 2011. Nonstructural carbohydrate content in the stem at full heading contributes to high performance of ripening in heat-tolerant rice cultivar Nikomaru. Crop Science, 51, 818–828.

Pan S G, Liu H D, Mo Z W, Patterson B, Duan M Y, Tian H, Xu S J, Tang X R. 2016. Effects of nitrogen and shading on root morphologies, nutrient accumulation, and photosynthetic parameters in different rice genotypes. Scientific Reports, 6, 32148.

Shao L P, Li G, Zhao Q N, Li Y B, Sun Y T, Wang W N, Cai C, Chen W P, Liu R H, Luo W H, Yin X Y, Lee X H. 2020. The fertilization effect of global dimming on crop yields is not attributed to an improved light interception. Global Change Biology, 26, 1697–1713.

Shi W J, Muthirajan R, Rahman H, Selvam J, Peng S B, Zou Y B, Jagadish K S V. 2013. Source–sink dynamics and proteomic reprogramming under elevated night temperature and their impact on rice yield and grain quality. New Phytologist, 197, 825–837.

Tong C, Chen Y L, Tang F F, Xu F F, Huang Y, Chen H, Bao J S. 2014. Genetic diversity of amylose content and RVA pasting parameters in 20 rice accessions grown in Hainan, China. Food Chemistry, 161, 239–245.

Toutounji M R, Farahnaky A, Santhakumar A B, Oli P, Butardo Jr V M, Blanchard C L. 2019. Intrinsic and extrinsic factors affecting rice starch digestibility. Trends in Food Science & Technology, 88, 10–22.

Tsukaguchi T, Tanaka R, Inoue H, Nakagawa H. 2018. Effects of high temperature and shading on grain abscisic acid content and grain filling pattern in rice (Oryza sativa L.). Plant Production Science, 21, 407–412.

Wang L, Deng F, Ren W J. 2015. Shading tolerance in rice is related to better light harvesting and use efficiency and grain filling rate during grain filling period. Field Crops Research, 180, 54–62.

Wang L, Deng F, Ren W J, Yang W Y. 2013. Effects of shading on starch pasting characteristics of indica hybrid rice (Oryza sativa L.). PLoS ONE, 8, e68220.

Wang C L, Zhang Y D, Zhu Z, Chen T, Zhao Q Y, Zhong W G, Yang J, Yao S, Zhou L H, Zhao L, Li Y S. 2017. Research progress on the breeding of japonica super rice varieties in Jiangsu Province, China. Journal of Integrative Agriculture, 16, 992–999.

Wang G Q, Hao S S, Gao B, Chen M X, Liu Y G, Yang J C, Ye N H, Zhang J H. 2017. Regulation of gene expression in the remobilization of carbon reserves in rice stems during grain filling. Plant & Cell Physiology, 58, 1391–1404.

Wang G Q, Zhang J H. 2020. Carbohydrate, hormone and enzyme regulations of rice grain filling under post-anthesis soil drying. Environmental and Experimental Botany, 178, 104165.

Wang Y J, Wei Y, Yan X T, Ge M J, Meng T Y, Zhang H C, Dai Q G, Huo Z Y, Xu K, Fei X R. 2014. Effects of light, nitrogen and their interaction on grain yield and nitrogen, phosphorus and potassium absorption in japonica super rice. Acta Agronomica Sinica, 40, 1235–1244. (in Chinese)

Wang Y Y, Meng Z, Lyu R, Huang G, He Q S, Cheng T T. 2020. Spatiotemporal changes of surface solar radiation: Implication for air pollution and rice yield in East China. Science of the Total Environment, 739, 140361.

Wei H Y, Zhu Y, Qiu S, Han C, Hu L, Xu D, Zhou N B, Xing Z P, Hu Y J, Cui P Y, Dai Q G, Zhang H C. 2018. Combined effect of shading time and nitrogen level on grain filling and grain quality in japonica super rice. Journal of Integrative Agriculture, 17, 2405–2417.

Wild M, Wacher S, Yang S, Sanchez-Lorenzo A. 2021. Evidence for clear-sky dimming and brightening in central Europe. Geophysical Research Letters, 48, e2020GL092216.

Wu L M, Zhang W J, Ding Y F, Zhang J W, Cambula E D, Weng F, Liu Z H, Ding C Q, Tang S, Chen L, Wang S H, Li G H. 2017. Shading contributes to the reduction of stem mechanical strength by decreasing cell wall synthesis in japonica rice (Oryza sativa L.). Frontiers in Plant Science, 8, 881.

Xie X B, Shan S L, Wang Y M, Cao F B, Chen J N, Huang M, Zou Y B. 2019. Dense planting with reducing nitrogen rate increased grain yield and nitrogen use efficiency in two hybrid rice varieties across two light conditions. Field Crops Research, 236, 24–32.

Xu H G, Zhong G R, Lin J J, Ding Y F, Li G H, Wang S H, Liu Z H, Tang S, Ding C Q. 2015. Effect of nitrogen management during the panicle stage in rice on the nitrogen utilization of rice and succeeding wheat crops. European Journal of Agronomy, 70, 41–47.

Xu Y J, Ying Y N, Ouyang S H, Duan X L, Sun H, Jiang S K, Sun S C, Bao J S. 2018. Factors affecting sensory quality of cooked japonica rice. Rice Science, 25, 330–339.

Yang J C, Zhang J H. 2010. Grain-filling problem in ‘super’ rice. Journal of Experimental Botany, 61, 1–5.

Yoshida H, Takehisa K, Kojima T, Ohno H, Sasaki K, Nakagawa H. 2016. Modeling the effects of N application on growth, yield and plant properties associated with the occurrence of chalky grains of rice. Plant Production Science, 19, 30–42.

Zhang J, Zhang Y Y, Song N Y, Chen Q L, Sun H Z, Peng T, Huang S, Zhao Q Z. 2021. Response of grain-filling rate and grain quality of mid-season indica rice to nitrogen application. Journal of Integrative Agriculture, 20, 1465–1473.

Zhen F X, Zhou J J, Mahmood A, Wang W, Chang X N, Liu B, Liu L L, Cao W X, Zhu Y, Tang L. 2020. Quantifying the effects of short-term heat stress at booting stage on nonstructural carbohydrates remobilization in rice. The Crop Journal, 8, 194–212.

Zhou H, Xia D, He Y Q. 2020. Rice grain quality - Traditional traits for high quality rice and health-plus substances. Molecular Breeding, 40, 1.

Zhu D W, Zhang H C, Guo B W, Xu K, Dai Q G, Wei H Y, Gao H, Hu Y J, Cui P Y, Huo Z Y. 2017. Effects of nitrogen level on yield and quality of japonica soft super rice. Journal of Integrative Agriculture, 16, 1018–1027.

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