中国农业科学 ›› 2023, Vol. 56 ›› Issue (2): 203-216.doi: 10.3864/j.issn.0578-1752.2023.02.001
陈吉浩1,2(),周界光1(),曲翔汝1,王素容1,唐华苹1,蒋云3,唐力为4,$\boxed{\hbox{兰秀锦}}$1,魏育明1,周景忠5(),马建1()
收稿日期:
2022-07-05
接受日期:
2022-08-12
出版日期:
2023-01-16
发布日期:
2023-02-07
通讯作者:
周景忠,E-mail:作者简介:
陈吉浩,E-mail:基金资助:
CHEN JiHao1,2(),ZHOU JieGuang1(),QU XiangRu1,WANG SuRong1,TANG HuaPing1,JIANG Yun3,TANG LiWei4,$\boxed{\hbox{LAN XiuJin}}$1,WEI YuMing1,ZHOU JingZhong5(),MA Jian1()
Received:
2022-07-05
Accepted:
2022-08-12
Online:
2023-01-16
Published:
2023-02-07
摘要:
【目的】挖掘小麦胚大小性状相关的数量性状位点,解析胚大小与其他重要农艺性状之间的相关性,为胚相关性状QTL的精细定位及育种利用奠定基础。【方法】以四倍体小麦矮兰麦(Ailanmai)和野生二粒小麦(LM001)构建的121份F8代重组自交系群体(AM群体)作为研究材料,将其分别种植于成都市崇州试验基地(2018、2019和2020年)、成都市温江区试验基地(2020年)和雅安市试验基地(2020年),调查5个环境下的胚长、胚宽、胚长/胚宽、胚长/粒长、胚宽/粒宽以及胚面积6个性状,结合基于小麦55K SNP芯片构建的遗传连锁图谱对上述6个性状进行QTL定位。【结果】胚大小性状呈近似正态分布,符合数量性状的遗传特征。QTL定位共检测到27个胚大小相关性状的QTL,其中,7个分别控制胚长和胚宽的QTL可解释7.75%—21.74%和7.67%—33.29%的表型变异,共检测到5个在多环境稳定表达的主效QTL:QEL.sicau-AM-3B、QEW.sicau-AM-2B、QEW/KW.sicau-AM-2B、QEL/EW.sicau-AM-2B-1和QEA.sicau-AM-2B,其贡献率分别为11.88%—21.74%、21.77%—33.29%、8.80%—24.92%、12.79%—31.13%和10.47%—20.67%。另外,上述胚相关的位点形成4个QTL簇,分别为1B控制胚长/粒长和胚长的QTL簇,2B控制胚宽、胚长/胚宽、胚宽/粒宽以及胚面积的QTL簇,3B控制胚长和胚面积的QTL簇,6B控制胚长/胚宽、胚宽/粒宽的QTL簇。确定胚大小与小麦籽粒大小具有显著正相关性,且发现胚长主效位点QEL.sicau-AM-3B与粒长主效位点共定位,但胚宽主效位点QEW.sicau-AM-2B独立于粒宽主效位点存在。在主效QTL所在物理区间鉴定获得4个可能参与胚大小调控的基因。【结论】鉴定到5个控制胚相关性状的稳定表达的主效QTL:QEL.sicau-AM-3B、QEW.sicau-AM-2B、QEW/KW.sicau-AM-2B、QEL/EW.sicau-AM-2B-1和QEA.sicau-AM-2B,其中,QEW.sicau-AM-2B可能为新的QTL。
陈吉浩, 周界光, 曲翔汝, 王素容, 唐华苹, 蒋云, 唐力为, $\boxed{\hbox{兰秀锦}}$, 魏育明, 周景忠, 马建. 四倍体小麦胚大小性状QTL定位与分析[J]. 中国农业科学, 2023, 56(2): 203-216.
CHEN JiHao, ZHOU JieGuang, QU XiangRu, WANG SuRong, TANG HuaPing, JIANG Yun, TANG LiWei, $\boxed{\hbox{LAN XiuJin}}$, WEI YuMing, ZHOU JingZhong, MA Jian. Mapping and Analysis of QTL for Embryo Size-Related Traits in Tetraploid Wheat[J]. Scientia Agricultura Sinica, 2023, 56(2): 203-216.
表1
AM群体亲本及其RIL胚相关性状表型变异"
性状 Trait | 环境 Environment | 亲本 Parents | 重组自交系群体 RIL population | |||||||
---|---|---|---|---|---|---|---|---|---|---|
Ailanmai | LM001 | 范围 Range | 平均值 Mean | 标准差 SD | 变异系数 CV | 偏度 Skewness | 峰度 Kurtosis | 广义遗传力 H2 | ||
EL (mm) | 2020WJ | 2.58 | 3.09a | 2.13—3.99 | 3.20 | 0.31 | 0.10 | -0.38 | 0.81 | 0.75 |
2020YA | 2.55 | 3.08a | 2.14—4.00 | 2.94 | 0.41 | 0.14 | 0.82 | 0.13 | ||
2020CZ | 2.51 | 3.08a | 2.29—3.99 | 3.06 | 0.36 | 0.12 | 0.37 | -0.22 | ||
2019CZ | 2.64 | 3.10a | 2.13—4.00 | 3.07 | 0.35 | 0.11 | 0.12 | 0.01 | ||
2018CZ | 2.43 | 2.75a | 2.43—4.00 | 3.13 | 0.31 | 0.10 | 0.21 | 0.04 | ||
BLUP | 2.62 | 3.02 | 2.45—3.37 | 3.00 | 0.17 | 0.06 | 0.59 | 0.81 | ||
EW (mm) | 2020WJ | 1.95a | 1.88 | 1.48—2.95 | 2.02 | 0.33 | 0.16 | 0.84 | 0.05 | 0.76 |
2020YA | 2.14a | 1.93 | 1.51—2.94 | 1.98 | 0.30 | 0.15 | 0.94 | 0.65 | ||
2020CZ | 1.98b | 1.91 | 1.40—3.00 | 2.08 | 0.36 | 0.17 | 0.66 | -0.12 | ||
2019CZ | 2.19a | 1.99 | 1.43—3.00 | 2.06 | 0.32 | 0.16 | 1.03 | 1.19 | ||
2018CZ | 1.90a | 1.54 | 1.45—3.00 | 2.01 | 0.30 | 0.15 | 1.16 | 1.48 | ||
BLUP | 2.02 | 1.86 | 1.64—3.00 | 1.95 | 0.14 | 0.07 | 0.14 | 0.01 | ||
EL/KL | 2020WJ | 0.39b | 0.36 | 0.31—0.49 | 0.38 | 0.03 | 0.08 | 0.85 | 0.64 | 0.37 |
2020YA | 0.38b | 0.36 | 0.27—0.41 | 0.34 | 0.03 | 0.09 | 0.12 | -0.29 | ||
2020CZ | 0.40b | 0.37 | 0.29—0.48 | 0.35 | 0.03 | 0.09 | 0.93 | 3.09 | ||
2019CZ | 0.40b | 0.36 | 0.29—0.49 | 0.36 | 0.03 | 0.08 | 0.93 | 2.12 | ||
2018CZ | 0.39a | 0.34 | 0.34—0.49 | 0.38 | 0.03 | 0.08 | 1.21 | 1.42 | ||
BLUP | 0.37 | 0.36 | 0.33—0.38 | 0.35 | 0.01 | 0.03 | -0.66 | 1.88 | ||
EW/KW | 2020WJ | 0.50b | 0.56 | 0.42—0.71 | 0.54 | 0.06 | 0.11 | 0.27 | -0.29 | 0.64 |
2020YA | 0.56 | 0.57 | 0.44—0.81 | 0.56 | 0.07 | 0.13 | 0.76 | 0.76 | ||
2020CZ | 0.50b | 0.57 | 0.39—0.76 | 0.57 | 0.07 | 0.12 | 0.18 | 0.23 | ||
2019CZ | 0.56 | 0.58 | 0.39—0.69 | 0.54 | 0.06 | 0.11 | 0.01 | -0.15 | ||
2018CZ | 0.49b | 0.46 | 0.42—0.65 | 0.55 | 0.06 | 0.11 | 0.26 | -0.19 | ||
BLUP | 0.53 | 0.55 | 0.47—0.65 | 0.55 | 0.03 | 0.05 | 0.22 | 0.14 | ||
EL/EW | 2020WJ | 1.33b | 1.65 | 1.20—1.99 | 1.65 | 0.16 | 0.10 | -0.41 | 0.16 | 0.72 |
2020YA | 1.20 | 1.60a | 1.15—1.99 | 1.49 | 0.16 | 0.11 | 0.71 | 0.68 | ||
2020CZ | 1.28 | 1.62a | 1.12—2.52 | 1.54 | 0.2 | 0.13 | 1.78 | 7.93 | ||
2019CZ | 1.23 | 1.56 | 1.16—2.14 | 1.53 | 0.17 | 0.11 | 0.37 | 0.92 | ||
2018CZ | 1.28b | 1.79 | 1.28—2.16 | 1.6 | 0.16 | 0.10 | 0.33 | 0.58 | ||
BLUP | 1.31 | 1.63 | 1.20—1.78 | 1.55 | 0.11 | 0.07 | -0.52 | 0.47 | ||
EA (mm2) | 2020WJ | 3.62a | 4.17 | 2.44—6.26 | 4.39 | 0.69 | 0.16 | 0.10 | -0.01 | 0.75 |
2020YA | 3.93a | 4.29 | 2.56—6.35 | 4.07 | 0.81 | 0.20 | 0.59 | -0.22 | ||
2020CZ | 3.56a | 4.24 | 2.65—6.18 | 4.27 | 0.72 | 0.17 | -0.05 | -0.16 | ||
2019CZ | 4.16a | 4.44 | 2.74—6.17 | 4.35 | 0.73 | 0.17 | 0.25 | -0.38 | ||
2018CZ | 3.33b | 3.04 | 2.76—6.63 | 4.33 | 0.71 | 0.16 | 0.39 | 0.32 | ||
BLUP | 3.80 | 4.07 | 3.22—5.48 | 4.22 | 0.46 | 0.11 | -0.12 | -0.21 |
表2
不同环境下胚相关性状的相关性分析"
性状 Trait | 环境 Environment | 2020WJ | 2020YA | 2020CZ | 2019CZ | 2018CZ | BLUP |
---|---|---|---|---|---|---|---|
胚长 EL | 2020WJ | 1 | |||||
2020YA | 0.34** | 1 | |||||
2020CZ | 0.49** | 0.38** | 1 | ||||
2019CZ | 0.37** | 0.32** | 0.48** | 1 | |||
2018CZ | 0.35** | 0.15** | 0.39** | 0.36** | 1 | ||
BLUP | 0.69** | 0.58** | 0.68** | 0.68** | 0.65** | 1 | |
胚宽 EW | 2020WJ | 1 | |||||
2020YA | 0.39** | 1 | |||||
2020CZ | 0.38** | 0.38** | 1 | ||||
2019CZ | 0.43** | 0.25** | 0.47** | 1 | |||
2018CZ | 0.36** | 0.41 | 0.34** | 0.26* | 1 | ||
BLUP | 0.63** | 0.66** | 0.65** | 0.60** | 0.59** | 1 | |
胚长/粒长 EL/KL | 2020WJ | 1 | |||||
2020YA | 0.06 | 1 | |||||
2020CZ | 0.46** | 0.19 | 1 | ||||
2019CZ | 0.45** | 0.08 | 0.43** | 1 | |||
2018CZ | 0.37** | 0.10 | 0.27* | 0.31** | 1 | ||
BLUP | 0.68** | 0.39** | 0.64** | 0.62** | 0.51** | 1 | |
胚宽/粒宽 EW/KW | 2020WJ | 1 | |||||
2020YA | 0.29** | 1 | |||||
2020CZ | 0.36** | 0.30** | 1 | ||||
2019CZ | 0.39** | 0.35** | 0.32** | 1 | |||
2018CZ | 0.54** | 0.35** | 0.42** | 0.32** | 1 | ||
BLUP | 0.72** | 0.69** | 0.72** | 0.68** | 0.69** | 1 | |
胚长/胚宽 EL/EW | 2020WJ | 1 | |||||
2020YA | 0.42** | 1 | |||||
2020CZ | 0.48** | 0.61** | 1 | ||||
2019CZ | 0.46** | 0.42** | 0.56** | 1 | |||
2018CZ | 0.63** | 0.44** | 0.43** | 0.37** | 1 | ||
BLUP | 0.78** | 0.78** | 0.73** | 0.71** | 0.73** | 1 | |
胚面积 EA | 2020WJ | 1 | |||||
2020YA | 0.44** | 1 | |||||
2020CZ | 0.62** | 0.37** | 1 | ||||
2019CZ | 0.58** | 0.32** | 0.48** | 1 | |||
2018CZ | 0.51** | 0.41** | 0.56** | 0.54** | 1 | ||
BLUP | 0.80** | 0.69** | 0.79** | 0.75** | 0.75** | 1 |
表3
胚与其他农艺性状的相关性"
性状 Trait | 分蘖数 Tiller number | 株高 Plant height | 粒长 Kernel length | 粒宽 Kernel width | 千粒重 Thousand-kernel weight | 开花期 Flowering period |
---|---|---|---|---|---|---|
胚长 EL | 0.03 | 0.62** | 0.71** | 0.33** | 0.71** | -0.11 |
胚宽 EW | -0.06 | 0.16 | 0.02 | 0.36** | 0.31** | 0.08 |
胚长/粒长 EL/KL | -0.27** | 0.08 | -0.19* | 0.36** | 0.08 | 0.11 |
胚宽/粒宽 EW/KW | -0.12 | -0.01 | -0.07 | -0.15 | 0.17 | 0.13 |
胚长/胚宽 EL/EW | 0.07 | 0.33** | 0.53** | -0.08 | 0.23* | -0.13 |
胚面积 EA | -0.02 | 0.42** | 0.38** | 0.40** | 0.57** | -0.02 |
表4
胚相关性状的QTL定位结果"
性状 Trait | 数量性状位点 QTL | 环境Environment | 染色体 Chromosome | 左标记 Left marker | 右标记 Right marker | 位置 Position (cM) | 阈值 LOD | 表型变异率 PVE (%) | 加性效应 Add |
---|---|---|---|---|---|---|---|---|---|
胚长 EL | QEL.sicau-AM-3B | 2020WJ | 3B | AX-110375013 | AX-110569582 | 6.91—7.34 | 5.15 | 16.21 | -0.11 |
2020YA | 3B | AX-110375013 | AX-110569582 | 6.91—7.34 | 3.63 | 11.88 | -0.10 | ||
2020CZ | 3B | AX-111112626 | AX-110375013 | 6.91—7.34 | 4.60 | 14.95 | -0.10 | ||
2019CZ | 3B | AX-110375013 | AX-110569582 | 6.91—7.34 | 4.90 | 18.99 | -0.12 | ||
BLUP | 3B | AX-110375013 | AX-110569582 | 6.91—7.34 | 8.41 | 21.74 | -0.09 | ||
QEL.sicau-AM-1B-1 | 2020CZ | 1B | AX-108913437 | AX-111510846 | 46.84—47.27 | 2.78 | 8.28 | 0.08 | |
BLUP | 1B | AX-108913437 | AX-111510846 | 46.84—47.27 | 4.22 | 9.77 | 0.06 | ||
QEL.sicau-AM-1B-2 | 2020YA | 1B | AX-111720572 | AX-110579523 | 50.30—51.15 | 4.14 | 13.37 | 0.11 | |
QEL.sicau-AM-7B | 2020WJ | 7B | AX-110995675 | AX-109951040 | 171.56—172.01 | 2.61 | 7.75 | 0.07 | |
胚宽 EW | QEW.sicau-AM-2B | 2020WJ | 2B | AX-109390001 | AX-110571221 | 0.00—1.92 | 7.42 | 24.13 | 0.10 |
2020CZ | 2B | AX-109390001 | AX-110571221 | 0.00—1.92 | 7.41 | 22.40 | 0.10 | ||
2019CZ | 2B | AX-109390001 | AX-110571221 | 0.00—1.92 | 6.19 | 21.77 | 0.09 | ||
2018CZ | 2B | AX-109390001 | AX-110571221 | 0.00—1.92 | 6.50 | 29.41 | 0.11 | ||
BLUP | 2B | AX-109390001 | AX-110571221 | 0.00—1.92 | 12.89 | 33.29 | 0.08 | ||
QEW.sicau-AM-2A-1 | 2020WJ | 2A | AX-109455008 | AX-109024104 | 70.71—74.53 | 2.55 | 7.67 | 0.06 | |
2019CZ | 2A | AX-109455008 | AX-109024104 | 70.71—74.53 | 2.75 | 9.12 | 0.06 | ||
BLUP | 2A | AX-109455008 | AX-109024104 | 70.71—74.53 | 4.80 | 10.74 | 0.05 | ||
QEW.sicau-AM-2A-2 | 2020CZ | 2A | AX-109934383 | AX-109314401 | 98.92—105.82 | 3.31 | 9.79 | 0.07 | |
胚长/粒长 EL/KL | QEL/KL.sicau-AM-1B-1 | 2020YA | 1B | AX-109379021 | AX-111720572 | 49.00—50.30 | 2.78 | 12.45 | 0.01 |
QEL/KL.sicau-AM-1B-2 | 2020CZ | 1B | AX-108913437 | AX-111510846 | 46.84—47.27 | 3.07 | 12.68 | 0.01 | |
QEL/KL.sicau-AM-4B | 2019CZ | 4B | AX-109052054 | AX-109931786 | 11.56—12.43 | 2.77 | 11.29 | 0.01 | |
QEL/KL.sicau-AM-5B | 2019CZ | 5B | AX-109990234 | AX-108886889 | 76.90—79.56 | 2.54 | 10.13 | -0.01 | |
QEL/KL.sicau-AM-2A | BLUP | 2A | AX-109455008 | AX-109024104 | 70.71—74.53 | 3.70 | 9.03 | 0.01 | |
QEL/KL.sicau-AM-2B | BLUP | 2B | AX-109390001 | AX-110571221 | 0.00—1.92 | 6.66 | 20.67 | 0.01 | |
胚宽/粒宽 EW/KW | QEW/KW.sicau-AM-2B | 2020CZ | 2B | AX-109390001 | AX-110571221 | 0.00—1.92 | 4.05 | 12.49 | 0.02 |
2019CZ | 2B | AX-109390001 | AX-110571221 | 0.00—1.92 | 2.57 | 8.80 | 0.02 | ||
2018CZ | 2B | AX-109390001 | AX-110571221 | 0.00—1.92 | 5.16 | 24.92 | 0.02 | ||
BLUP | 2B | AX-109390001 | AX-110571221 | 0.00—1.92 | 6.65 | 20.68 | 0.01 | ||
QEW/KW.sicau-AM-4A | 2019CZ | 4A | AX-109272723 | AX-110524110 | 11.88—31.77 | 2.78 | 13.49 | -0.02 | |
QEW/KW.sicau-AM-6B | 2020CZ | 6B | AX-111012548 | AX-108984460 | 136.24—162.46 | 3.47 | 10.94 | -0.02 | |
胚长/胚宽 EL/EW | QEL/EW.sicau-AM-2B-1 | 2020WJ | 2B | AX-109390001 | AX-110571221 | 0.00—1.92 | 8.00 | 21.55 | -0.08 |
2020YA | 2B | AX-109390001 | AX-110571221 | 0.00—1.92 | 4.04 | 16.41 | -0.09 | ||
2020CZ | 2B | AX-109390001 | AX-110571221 | 0.00—1.92 | 8.83 | 25.69 | -0.08 | ||
2019CZ | 2B | AX-109390001 | AX-110571221 | 0.00—1.92 | 4.19 | 12.79 | -0.06 | ||
2018CZ | 2B | AX-109390001 | AX-110571221 | 0.00—1.92 | 5.45 | 23.41 | -0.07 | ||
BLUP | 2B | AX-109390001 | AX-110571221 | 0.00—1.92 | 11.64 | 31.13 | -0.06 | ||
性状 Trait | 数量性状位点 QTL | 环境Environment | 染色体 Chromosome | 左标记 Left marker | 右标记 Right marker | 位置 Position (cM) | 阈值 LOD | 表型变异率 PVE (%) | 加性效应 Add |
QEL/EW.sicau-AM-2B-2 | 2018CZ | 2B | AX-111525915 | AX-109297362 | 64.00—66.68 | 3.01 | 12.12 | -0.05 | |
QEL/EW.sicau-AM-2B-3 | 2020WJ | 2B | AX-110447950 | AX-108942420 | 33.21—39.07 | 3.03 | 7.89 | -0.05 | |
QEL/EW.sicau-AM-6B | 2020CZ | 6B | AX-111012548 | AX-108984460 | 136.24—162.46 | 3.38 | 9.10 | 0.05 | |
QEL/EW.sicau-AM-2A-1 | 2020WJ | 2A | AX-109314401 | AX-110069674 | 105.82—124.56 | 3.14 | 8.05 | -0.05 | |
QEL/EW.sicau-AM-2A-2 | BLUP | 2A | AX-110492886 | AX-109523500 | 92.49—95.29 | 2.76 | 6.33 | -0.03 | |
胚面积 EA | QEA.sicau-AM-2B | 2020WJ | 2B | AX-109390001 | AX-110571221 | 0.00—1.92 | 4.23 | 11.34 | 0.25 |
2019CZ | 2B | AX-109390001 | AX-110571221 | 0.00—1.92 | 2.88 | 10.47 | 0.21 | ||
2018CZ | 2B | AX-109390001 | AX-110571221 | 0.00—1.92 | 3.01 | 15.22 | 0.25 | ||
BLUP | 2B | AX-109390001 | AX-110571221 | 0.00—1.92 | 6.66 | 20.67 | 0.01 | ||
QEA.sicau-AM-2A-1 | 2020WJ | 2A | AX-109437513 | AX-109974497 | 66.15—67.06 | 4.29 | 11.41 | 0.25 | |
QEA.sicau-AM-2A-2 | BLUP | 2A | AX-109455008 | AX-109024104 | 70.71—74.53 | 3.07 | 9.03 | 0.01 | |
QEA.sicau-AM-3B | 2019CZ | 3B | AX-110375013 | AX-110569582 | 6.91—7.34 | 4.65 | 17.65 | -0.28 | |
QEA.sicau-AM-4B | 2020CZ | 4B | AX-108737791 | AX-110544397 | 20.75—22.08 | 3.47 | 13.18 | 0.23 |
表5
胚相关性状QTL的物理区间"
性状 Trait | QTL | 遗传标记区间 Flanking markers | 野生二粒小麦参考基因组 v2.0物理位置 Wild Emmer v2.0 physical map (Mb) | 中国春参考基因组v2.1物理位置 Chines Spring wheat v2.1 physical map (Mb) |
---|---|---|---|---|
胚长 EL | QEL.sicau-AM-3B | AX-110375013—AX-110569582 | 695.42—696.83 | 690.52—692.06 |
QEW.sicau-AM-2B | AX-109390001—AX-110571221 | 6.41—53.55 | 3.69—9.15 | |
胚宽 EW | QEL/KL.sicau-AM-1B-1 | AX-108913437—AX-111510846 | 409.17—411.07 | 395.61—397.54 |
QEL.sicau-AM-1B-2 | AX-111720572—AX-110579523 | 432.54—447.52 | 418.79—434.50 | |
胚长/粒长 EL/KL | QEW.sicau-AM-2A-1 | AX-109455008—AX-109024104 | 614.32—618.45 | 615.12—619.25 |
胚宽/粒宽 EW/KW | QEW/KW.sicau-AM-2B | AX-109390001—AX-110571221 | 6.41—53.55 | 3.69—9.15 |
QEA.sicau-AM-2B | AX-109390001—AX-110571221 | 6.41—53.55 | 3.69—9.15 | |
胚面积 EA | QEW/KW.sicau-AM-6B | AX-111012548—AX-108984460 | 712.45—723.67 | 721.64—730.61 |
表6
QEW.sicau-AM-2B、QEL.sicau-AM-3B与前人报道的籽粒相关QTL和分子标记"
性状 Trait | QTL | 染色体Chromosome | 左标记 Left marker | 右标记 Right marker | ||
---|---|---|---|---|---|---|
标记ID Marker ID | 物理位置 WE v2.0/CS 2.1 (Mb) | 标记ID Marker ID | 物理位置 WE v2.0/CS 2.1(Mb) | |||
粒长Kernel length | qKL.3B | 3B | * | * | Xwmc754 | 21.66/14.4 |
QGl.ccsu-3B.1 | 3B | Xgwm566 | 91.51/77.7 | Xgwm376 | 64.63/38.9 | |
QKL.sicau-AM-3B | 3B | AX-110375013 | 695.42/690.52 | AX-111112626 | 675.62/655.4 | |
胚长EL | QEL.sicau-AM-3B | 3B | AX-110375013 | 695.42/690.52 | AX-110569582 | 696.83/692.06 |
胚宽EW | QEW.sicau-AM-2B | 2B | AX-109390001 | 6.41/3.69 | AX-110571221 | 53.55/9.15 |
千粒重 Thousand grain weight | QTgw.crc-2B | 2B | Xwmc661 | 6.49/9.53 | * | * |
粒重Kernel weight | qKLW2B-1 | 2B | Xgwm501 | 98.15/680.09 | Xmag3512 | 780.69/771.06 |
粒宽Kernel width | QKw.ncl-2B.1 | 2B | Xbarc183 | 123.83/2.60 | Xbarc13 | 134.83/125.20 |
QTkw.ncl-2B.1 | 2B | Xbarc183 | 123.83/2.60 | Xbarc7 | 134.82/125.20 |
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