Bose J, Rodrigo-Moreno A, Shabala S. 2014. ROS homeostasis in halophytes in the context of salinity stress tolerance. Journal of Experimental Botany, 65, 1241–1257.

Choudhury F K, Rivero R M, Blumwald E, Mittler R. 2017. Reactive oxygen species, abiotic stress and stress combination. The Plant Journal, 90, 856–867.

Darabi M, Seddigh S, Abarshahr M. 2017. Structural, functional, and phylogenetic studies of cytochrome P450 (CYP) enzyme in seed plants by bioinformatics tools. Caryologia, 70, 62–76.

Daszkowska-Golec A. 2016. The role of abscisic acid in drought stress: How ABA helps plants to cope with drought stress. Drought Stress Tolerance in Plants, 2, 123–151.

Fujita Y, Fujita M, Satoh R, Maruyama K, Parvez M M, Seki M, Hiratsu K, Ohme-Takagi M, Shinozaki K, Yamaguchi-Shinozaki K. 2005. AREB1 is a transcription activator of novel ABRE-dependent ABA signaling that enhances drought stress tolerance in Arabidopsis. The Plant Cell, 17, 3470–3488.

Fujita Y, Yoshida T, Yamaguchi-Shinozaki K. 2013. Pivotal role of the AREB/ABF-SnRK2 pathway in ABRE-mediated transcription in response to osmotic stress in plants. Physiologia Plantarum, 147, 15–27.

Gao X R, Zhang H, Li X, Bai Y W, Peng K, Wang Z, Dai Z R, Bian X F, Zhang Q, Jia L C, Li Y, Liu Q C, Zhai H, Gao S P, Zhao N, He S Z. 2023. The B-box transcription factor IbBBX29 regulates leaf development and flavonoid biosynthesis in sweet potato. Plant Physiology, 191, 496–514.

Gong Z Z, Xiong L M, Shi H Z, Yang S H, Herrera-Estrella L R, Xu G H, Chao D Y, Li J R, Wang P Y, Qin F, Li J J, Ding Y L, Shi Y T, Wang Y, Yang Y Q, Guo Y, Zhu J K. 2020. Plant abiotic stress response and nutrient use efficiency. Science China (Life Sciences), 63, 635–674.

Hu L Y, Yue H, Zhang J Y, Li Y T S, Gong X Q, Zhou K, Ma F W. 2022. Overexpression of MdMIPS1 enhances drought tolerance and water-use efficiency in apple. Journal of Integrative Agriculture, 21, 1968–1981.

Jiang C J, Aono M, Tamaoki M, Maeda S, Sugano S, Mori M, Takatsuji H. 2008. SAZ, a new SUPERMAN-like protein, negatively regulates a subset of ABA-responsive genes in Arabidopsis. Molecular Genetics and Genomics, 279, 183–192.

Jiang X Q, Zhang C Q, Lü P T, Jiang G M, Liu X W, Dai F W, Gao J P. 2014. RhNAC3, a stress-associated NAC transcription factor, has a role in dehydration tolerance through regulatingosmotic stress-related genes in rose petals. Plant Biotechnology Journal, 12, 38–48.

Kang C, Zhai H, He S Z, Zhao N, Liu Q C. 2019. A novel sweetpotato bZIP transcription factor gene, IbbZIP1, is involved in salt and drought tolerance in transgenic Arabidopsis. Plant Cell Reports, 38, 1373–1382.

Kim Y Y, Jung K W, Yoo K S, Jeung J U, Shin J S. 2011. A stress-responsive caleosin-like protein, AtCLO4, acts as a negative regulator of ABA responses in Arabidopsis. Plant and Cell Physiology, 52, 874–884.

Kumar S, Stecher G, Li M, Knyaz C, Tamura K. 2018. MEGA X: Molecular evolutionary genetics analysis across computing platforms. Molecular Biology and Evolution, 35, 1547–1549.

Lescot M, Dehais P, Thijs G, Marchal K, Moreau Y, Van De Peer Y, Rouze P, Rombauts S. 2002. PlantCARE, a database of plant cis-acting regulatory elements and a portal to tools for in silico analysis of promoter sequences. Nucleic Acids Research, 30, 325–327.

Li R J, Zhai H, He S Z, Zhang H, Zhao N, Liu Q C. 2022. A geranylgeranyl pyrophosphate synthase gene, IbGGPS, increases carotenoid contents in transgenic sweetpotato. Journal of Integrative Agriculture, 21, 2538–2546.

Li Y, Zhang H, Zhang Q, Liu Q C, Zhai H, Zhao N, He S Z. 2019. An AP2/ERF gene, IbRAP2–12, from sweetpotato is involved in salt and drought tolerance in transgenic Arabidopsis. Plant Science, 281, 19–30.

Liang C Z, Meng Z H, Meng Z G, Malik W, Yan R, Lwin K M, Lin F Z, Wang Y, Sun G Q, Zhou T, Zhu T, Li J Y, Jin S X, Guo S D, Zhang R. 2016. GhABF2, a bZIP transcription factor, confers drought and salinity tolerance in cotton (Gossypium hirsutum L.). Scientific Reports, 6, 35040.

Liu D G, He S Z, Zhai H, Wang L J, Zhao Y, Wang B, Li R J, Liu Q C. 2014. Overexpression of IbP5CRenhances salt tolerance in transgenic sweetpotato. Plant Cell Tissue and Organ Culture, 117, 1–16.

Liu Q C. 2017. Improvement for agronomically important traits by gene engineering in sweetpotato. Breeding Science, 67, 15–26.

Liu Q C, Zhai H, Wang Y, Zhang D P. 2001. Efficient plantregeneration from embryogenic suspension cultures of sweetpotato. In Vitro Cellular & Developmental Biology (Plant), 37, 564–567.

Liu R D, Shen Y H, Wang M X, Liu R H, Cui Z Q, Li P Z, Wu Q D, Shen Q, Chen J, Zhang S P, Liu S D, Ma H J, Pang C Y, Ge C W. 2023. GhMYB102 promotes drought resistance by regulating drought-responsive genes and ABA biosynthesis in cotton (Gossypium hirsutum L.). Plant Science, 329, 111608.

Lu S N, Wang J Y, Chitsaz F, Derbyshire M K, Geer R C, Gonzales N R, Gwadz M, Hurwitz D I, Marchler G H, Song J S, Thanki N, Yamashita R A, Yang M Z, Zhang D C, Zheng C J, Lanczycki C J, Marchler-Bauer A. 2020. CDD/SPARCLE: the conserved domain database in 2020. Nucleic Acids Research, 48, D265–D268.

Miller G, Suzuki N, Ciftci-Yilmaz S, Mittler R. 2010. Reactive oxygen species homeostasis and signalling during drought and salinity stresses. Plant Cell and Environment, 33, 453–467.

Munemasa S, Hauser F, Park J, Waadt R, Brandt B, Schroeder J I. 2015. Mechanisms of abscisic acid-mediated control of stomatal aperture. Current Opinion in Plant Biology, 28, 154–162.

Pei D, Hua D P, Deng J P, Wang Z F, Song C P, Wang Y, Wang Y, Qi J S, Kollist H, Yang S H, Guo Y, Gong Z Z. 2022. Phosphorylation of the plasma membrane H+-ATPase AHA2 by BAK1 is required for ABA-induced stomatal closure in Arabidopsis. The Plant Cell, 34, 2708–2729.

Qin H, Wang J, Chen X B, Wang F F, Peng P, Zhou Y, Miao Y C, Zhang Y O, Gao Y D, Qi Y D, Zhou J H, Huang R F. 2019. Rice OsDOF15 contributes to ethylene-inhibited primary root elongation under salt stress. New Phytologist, 223, 798–813.

Qin T, Tian Q Z, Wang G F, Xiong L M. 2019. LOWER TEMPERATURE 1 enhances ABA responses and plant drought tolerance by modulating the stability and localization of C2-domain ABA-related proteins in Arabidopsis. Molecular Plant, 12, 1243–1258.

Ren Z T, He S Z, Zhou Y Y, Zhao N, Jiang T, Zhai H, Liu Q C. 2020. A sucrose non-fermenting–1-related protein kinase–1 gene, IbSnRK1, confers salt, drought and cold tolerance in sweet potato. The Crop Journal, 8, 905–917.

Rogers S O, Bendich A J. 1985. Extraction of DNA from milligram amounts of fresh, herbarium and mummified plant tissues. Plant Molecular Biology, 5, 69–76.

Sah S K, Jumaa S, Li J X, Reddy K R. 2022. Proteomic analysis response of rice (Oryza sativa) leaves to ultraviolet-B radiation stress. Frontiers in Plant Science, 13, 871331.

Salehi-Lisar S Y, Bakhshayeshan-Agdam H. 2020. Agronomic crop responses and tolerance to drought stress. Agronomic Crops, 3, 63–91.

Sun S F, Li X, Gao S P, Nie N, Zhang H, Yang Y F, He S Z, Liu Q C, Zhai H. 2022. A novel WRKY transcription factor from Ipomoea trifida, ItfWRKY70, confers drought tolerance in sweet potato. International Journal of Molecular Sciences, 23, 686.

Takahashi Y, Ebisu Y, Shimazaki K. 2017. Reconstitution of abscisic acid signaling from the receptor to DNA via bHLH transcription factors. Plant Physiology, 174, 815–822.

Tuteja N. 2007. Abscisic acid and abiotic stress signaling. Plant Signaling & Behavior, 2, 135–138.

Wang A M, Zhu M K, Luo Y H, Liu Y J, Li R S, Kou M, Wang X, Zhang Y G, Meng X Q, Zheng Y L, Ma D F. 2017. A sweet potato cinnamate 4-hydroxylase gene, IbC4H, increases phenolics content and enhances drought tolerance in tobacco. Acta Physiologiae Plantarum, 39, 276.

Wu X W, Liang Y, Gao H B, Wang J Y, Zhao Y, Hua L K, Yuan Y D, Wang A H, Zhang X H, Liu J F, Zhou J, Meng X B, Zhang D H, Lin S Y, Huang X H, Han B, Li J Y, Wang Y H. 2021. Enhancing rice grain production by manipulating the naturally evolved cis-regulatoryelement-containing inverted repeat sequence of OsREM20. Molecular Plant, 14, 997–1011.

Xing S H, Li R J, Zhao H Q, Zhai H, He S Z, Zhang H, Zhou Y Y, Zhao N, Gao S P, Liu Q C. 2023. The transcription factor IbNAC29 positively regulates the carotenoid accumulation in sweet potato. Horticulture Research, 10, uhad010.

Xue L Y, Wei Z H, Zhai H, Xing S H, Wang Y X, He S Z, Gao S P, Zhao N, Zhang H, Liu Q C. 2022. The IbPYL8-IbbHLH66-IbbHLH118 complex mediates the abscisic acid-dependent drought response in sweet potato. New Phytologist, 236, 2151–2171.

Yuan X Y, Xu S P, Wang M F, Lei Z H, Cui B. 2019. Cloning of PhTSJT1 gene from Phalaenopsis and its expression under low temperature stress. Genomics and Applied Biology, 38, 707–713.

Zhang H, Gao X R, Zhi Y H, Li X, Zhang Q, Niu J B, Wang J, Zhai H, Zhao N, Li J G, Liu Q C, He S Z. 2019. A non-tandem CCCH-type zinc-finger protein, IbC3H18, functions as a nuclear transcriptional activator and enhances abiotic stress tolerance in sweet potato. New Phytologist, 223, 1918–1936.

Zhang H, Wang Z, Li X, Gao X R, Dai Z R, Cui Y F, Zhi Y H, Liu Q C, Zhai H, Gao S P, Zhao N, He S Z. 2022. The IbBBX24-IbTOE3-IbPRX17 module enhances abiotic stress tolerance by scavenging reactive oxygen species in sweet potato. New Phytologist, 233, 1133–1152.

Zhang H, Zhang Q, Zhai H, Gao S P, Yang L, Wang Z, Xu Y T, Huo J X, Ren Z T, Zhao N, Wang X F, Li J G, Liu Q C, He S Z. 2020. IbBBX24 promotes the jasmonic acid pathway and enhances fusarium wilt resistance in sweet potato. The Plant Cell, 32, 1102–1123.

Zhang L, Yu Z P, Xu Y, Yu M, Ren Y, Zhang S Z, Yang G D, Huang J G, Yan K, Zheng C C, Wu C G. 2021. Regulation of the stability and ABA import activity of NRT1.2/NPF4.6 by CEPR2-mediated phosphorylation in Arabidopsis. Molecular Plant, 14, 633–646.

Zhang Y, Li J J, Li C R, Chen S K, Tang Q L, Xiao Y F, Zhong L X, Chen Y Y, Chen B W. 2022. Gene expression programs during callus development in tissue culture of two Eucalyptus species. BMC Plant Biology, 22, 1.

Zhao G Y, Cheng Q, Zhao Y T, Wu F F, Mu B B, Gao J P, Yang L, Yan J L, Zhang H F, Cui X, Chen Q Q, Lu F X, Ao Q Q, Amdouni A, Jiang Y Q, Yang B. 2023. The abscisic acid-responsive element binding factors MAPKKK18 module regulates abscisic acid-induced leaf senescence in Arabidopsis. The Journal of Biological Chemistry, 299, 103060.

Zhao L, Hu Y, Chong K, Wang T. 2010. ARAG1, an ABA-responsive DREB gene, plays a role in seed germination and drought tolerance of rice. Annals of Botany, 105, 401–409.

Zhao Y J, Zhang Y Y, Bai X Y, Lin R Z, Shi G Q, Du P P, Xiao K. 2022. TaNF-YB11, a gene of NF-Y transcription factor family in Triticum aestivum, confers drought tolerance on plants via modulating osmolyte accumulation and reactive oxygen species homeostasis. Journal of Integrative Agriculture, 21, 3114–3130.

Zhou Y Y, Zhai H, He S Z, Zhu H, Gao S P, Xing S H, Wei Z H, Zhao N, Liu Q C. 2020. The sweetpotato BTB-TAZ protein gene, IbBT4, enhances drought tolerance in transgenic Arabidopsis. Frontiers in Plant Science, 11, 877.

Zhou Y Y, Zhai H, Xing S H, Wei Z H, He S Z, Zhang H, Gao S P, Zhao N, Liu Q C. 2022. A novel small open reading frame gene, IbEGF, enhances drought tolerance in transgenic sweet potato. Frontiers in Plant Science, 13, 965069.

Zhou Y Y, Zhu H, He S Z, Zhai H, Zhao N, Xing S H, Wei Z H, Liu Q C. 2019. A novel sweetpotato transcription factor gene IbMYB116 enhances drought tolerance in transgenic Arabidopsis. Frontiers in Plant Science, 10, 1025.

Zhu H, Yang X, Wang X, Li Q Y, Guo J Y, Ma T, Zhao C M, Tang Y Y, Qiao L X, Wang J S, Sui J M. 2021. The sweetpotato beta-amylase gene IbBAM1.1 enhances drought and salt stress resistance by regulating ROS homeostasis and osmotic balance. Plant Physiology and Biochemistry, 168, 167–176.

Zhu H, Zhai H, He S Z, Zhang H, Gao S P, Liu Q C. 2022. A novel sweetpotato GATA transcription factor, IbGATA24, interacting with IbCOP9-5a positively regulates drought and salt tolerance. Environmental and Experimental Botany, 194, 104735.

Zhu H, Zhou Y Y, Zhai H, He S Z, Zhao N, Liu Q C. 2020. A novel sweetpotato WRKY transcription factor, IbWRKY2, positively regulates drought and salt tolerance in transgenic Arabidopsis. Biomolecules, 10, 506.

Zhu J K. 2002. Salt and drought stress signal transduction in plants. Annual Review of Plant Biology, 53, 247–273.

Zhu J K. 2016. Abiotic stress signaling and responses in plants. Cell, 167, 313–324.