Babar S, Baloch A, Qasim M, Wang J Y, Wang X L, Li Y X, Khalid S, Jiang C C. 2024. Unearthing the soil-bacteria nexus to enhance potassium bioavailability for global sustainable agriculture: A mechanistic preview. Microbiological Research, 288, 127885.

Biassoni M M, Vivas H, Guti´errez-Boem F H, Salvagiotti F. 2023. Changes in soil phosphorus (P) fractions and P bioavailability after 10 years of continuous P fertilization. Soil and Tillage Research, 232, 10577.  

Bremner J M. 1965. Total nitrogen. In: Methods of Soil Analysis. Part 2. Chemical and Microbiological Properties. American Society of Agronomy, USA. pp.1149–1178.

Brödlin D, Kaiser K, Kessler A, Hagedorn F. 2019. Drying and rewetting foster phosphorus depletion of forest soils. Soil Biology and Biochemistry, 128, 22–34.

Cao N, Zhi M L, Zhao W Q, Pang J Y, Hu W, Zhou Z G, Meng Y L. 2022. Straw retention combined with phosphorus fertilizer promotes soil phosphorus availability by enhancing soil P-related enzymes and the abundance of phoC and phoD genes. Soil & Tillage Research, 220, 105390.

Chai Y W, Chai Q, Han F X, Li Y W, Ma J T, Li R, Cheng H B, Chang L, Chai S X. 2022. Increasing yields while reducing soil nutrient accumulation by straw strip mulching in the dryland wheat (Triticum aestivum L.) cropping system of Northwest China. Agriculture, Ecosystems and Environment, 326, 107797.

Chan H L, Lam S K, Chen D L, Tang C X, Chen Q L, Roessner U, Hu H W. 2025. Nitrogen deficiency drives fungal compositional shifts without functional changes in wheat rhizosphere. Rhizosphere, 33, 101030.

Channab B E, Tayi F, Aqlil M, Akil A, Essamlali Y, Chakir A, Zahouily M. 2024. Graphene oxide, starch, and kraft lignin bio-nanocomposite controlled-release phosphorus fertilizer: Effect on P management and maize growth. International Journal of Biological Macromolecules, 282, 137190.

Chen X D, Wang Y, Guo B, Liu J L, Qiu G Y, Yakov K, Li H. 2025. Long-term green manuring increases phosphorus availability in soil by modifying microbial functional genes and enzyme activities. Applied Soil Ecology, 213, 106284.

Chen Y M, Xu X, Jiao X G, Sui Y Y, Liu X B, Zhang J Y, Zhou K, Zhang J M. 2018. Responses of labile organic nitrogen fractions and enzyme activities in eroded mollisols after 8-year manure amendment. Scientific Reports, 8, 14179.

Cheng L, Cui H, Ou Y, Wang L X, Bao M W. 2024. Redistribution of phosphorus fraction driven by organic carbon and microbial community during composting. Journal of Environmental Management, 370, 123003.

Chuan L M, He P, Jin J Y, Li S T, Grant C, Xu X P, Qiu S J, Zhao S C, Zhou W. 2013. Estimating nutrient uptake requirements for wheat in China. Field Crops Research, 146, 96–104.

Cross A F, Schlesinger W H. 1995. A literature review and evaluation of the Hedley fractionation: Applications to the biogeochemical cycle of soil phosphorus in natural ecosystems. Geoderma, 64, 197–214. 

Dai J, Bean B, Brown B, Bruening W, Edwards J, Flowers M, Karow R, Lee C, Morgan G, Ottman M, Ransom J, Wiersma J. 2016. Harvest index and straw yield of five classes of wheat. Biomass and Bioenergy, 85, 223–227.

Ding J P, Li J H, Liu J H, Zhang W F, Jia X P. 2022. ICT-based agricultural advisory services and nitrogen management practices: A case study of wheat production in China. Journal of Integrative Agriculture, 21, 1799–1811.

Dou Y X, Zhao H B, Yang H M, Wang T, Liu G F, Wang Z H, Malhi S. 2024. The first factor affecting dryland winter wheat grain yield under various mulching measures: Spike number. Journal of Integrative Agriculture, 23, 836–848.

Gao X L, Li X G, Zhao L, Kuzyakov Y. 2021. Shrubs magnify soil phosphorus depletion in Tibetan meadows: Conclusions from C: N:P stoichiometry and deep soil profiles. Science of the Total Environment, 785, 147320.

He P, Yang L P, Xu X P, Zhao S C, Chen F, Li S T, Tu S H, Jin J Y, Johnston A M. 2015. Temporal and spatial variation of soil available potassium in China (1990–2012). Field Crops Research, 173, 49–56.

Hedley M J, Stewart J W B, Chauhan B. 1982. Changes in inorganic and organic soil phosphorus fractions induced by cultivation practices and by laboratory incubations. Soil Science Society of America Journal, 46, 970–976.

Hinsinger P, Plassard C, Tang C, Jaillard B. 2003. Origins of root-mediated pH changes in the rhizosphere and their responses to environmental constraints: A review. Plant Soil, 248, 43–59.

Huo W G, Chai X F, Wang X H, William D B, Kafle A, Gu F. 2022. Indigenous arbuscular mycorrhizal fungi play a role in phosphorus depletion in organic manure amended high fertility soil. Journal of Integrative Agriculture, 21, 3051–3066.

Jadav K, Durgam M, Perli M, Mailapalli D R. 2024. Phosphorus transport modeling in soils treated with nano-phosphorus fertilizers. Plant Nano Biology, 10, 100112.

Jiang H M, Li S S, Zhang J F, Yang J C, Li L L, Zhang S Q, Guo J M, Liu L, Xie Y Q, Wang F Y. 2014. Transformation of external chemical nitrogen in soil organic nitrogen fractions and their relationship. Journal of Plant Nutrition and Fertilizer, 20, 1421–1430. (in Chinese)

Jin Z X, Wang J F, Chen J G, Zhang R X, Li Y, Lu Y T, He K K. 2020. Identifying the sources of nitrate in a small watershed using δ¹⁵N-δ¹⁸O isotopes of nitrate in the Kelan Reservoir, Guangxi, China. Agriculture, Ecosystems and Environment, 297, 106936.

Khan A, Yang X Y, Sun B H, Zhang S L, He B H. 2023. Responses of crop and soil phosphorus fractions to long-term fertilization regimes in a loess soil in northwest China. Agronomy, 13, 3072.

Li C, Ma X, Wang Y, Sun Q, Chen M S, Zhang C S, Ding S M, Dai Z H. 2024. Root-mediated acidification, phosphatase activity and the phosphorus-cycling microbial community enhance phosphorus mobilization in the rhizosphere of wetland plants. Water Research, 255, 121548.

Li T L, Xie Y H, Gao Z Q, Hong J P, Li L, Meng H S, Ma H M, Jia J X. 2019.Year-round film mulching system with monitored fertilization management improve grain yield and water and nitrogen use efficiencies of winter wheat in the dryland of the Loess Plateau, China. Environmental Science and Pollution Research, 26, 9524–9535.

Li Z L, Zhang W T, Qiu L X, Pan T L, Zheng W K, Kong B, Wang H L, Li C L, Liu Z G, Zhang M. 2021. Physiological-biochemical responses of wheat to blending controlled-release potassium chloride and soluble potassium chloride. Soil & Tillage Research, 212, 105058.

Liu J, Sui P, Cade-Menun B J, Hu Y F, Yang J J, Huang S M, Ma Y B. 2019. Molecular-level understanding of phosphorus transformation with long-term phosphorus addition and depletion in an alkaline soil. Geoderma, 353, 116–124.

Liu Y, Gao J P, Zhong M, Chen L Q, Zhang W Z. 2024. Effects of phosphorus and potassium supply on photosynthetic nitrogen metabolism, nitrogen absorption, and nitrogen utilization of hydroponic rice. Agronomy, 14, 1726.

Liu Z J, Yuan X H, Zhang Z X, Yang X Y, Ai C, Wang Z H, Fan Q L, Xu X P. 2025. Revisiting potassium-induced impacts on crop production and soil fertility based on thirty-three Chinese long-term experiments. Field Crops Research, 322, 109732.

Liu Z P, Shao M A, Wang Y Q. 2013. Spatial patterns of soil total nitrogen and soil total phosphorus across the entire Loess Plateau region of China. Geoderma, 197–198, 67–78.

Lopez G, Ahmadi S H, Amelung W, Athmann M, Ewert F, Gaiser T, Gocke M I, Kautz T, Postma J, Rachmilevitch S, Schaaf G, Schnepf A, Stoschus A, Watt M, Yu P, Seidel S J. 2023. Nutrient deficiency effects on root architecture and root-to-shoot ratio in arable crops. Frontier in Plant Science, 13, 1067498.

Muqaddas B, Lewis T, Esfandbod M, Chen C R. 2019. Responses of labile soil organic carbon and nitrogen pools to long-term prescribed burning regimes in a wet sclerophyll forest of southeast Queensland, Australia. Science of the Total Environment, 647, 110–120.

Quan Z, Huang B, Lu C Y, Shi Y, Chen X, Zhou J B, Fang Y T. 2018. Formation of extractable organic nitrogen in an agricultural soil: A ¹⁵N labeling study. Soil Biology and Biochemistry, 118, 161–165.

Ros G H, Hoffland E, Temminghoff E J M. 2010. Dynamics of dissolved and extractable organic nitrogen upon soil amendment with crop residues. Soil Biology and Biochemistry, 42, 2094–2101.

Shao W Y, Wang Q Z, Guan Q Y, Luo H P, Ma Y R, Zhang J. 2022. Distribution of soil available nutrients and their response to environmental factors based on path analysis model in arid and semi-arid area of northwest China. Science of the Total Environment, 827, 154254.

Sequeira C H, Alley M M, Jones B P. 2011. Evaluation of potentially labile soil organic carbon and nitrogen fractionation procedures. Soil Biology & Biochemistry, 43, 438–444.

Tiessen H J W B, Stewart J W B, Cole C V. 1984. Pathways of phosphorus transformations in soils of differing pedogenesis. Soil Science Society of America Journal, 48, 853–858.

Wan S Z, Lin G G, Liu B, Ding Y D, Li S L, Mao R. 2022. Contrasting responses of soil phosphorus pool and bioavailability to alder expansion in a boreal peatland, Northeast China. Catena, 212, 106128.

Wang C W, Xie Y K, Tan Z X. 2024. Soil potassium depletion in global cereal croplands and its implications. Science of the Total Environment, 907, 167875.

Wang H Y, Zhou J M, Du C W, Chen X Q. 2010. Potassium fractions in soils as affected by monocalcium phosphate, ammonium sulfate, and potassium chloride application. Pedosphere, 20, 368–377.

Wang M J, Feng X Q, Zhou Z Y, Ma H Y, Ge T D, Tang C X, Wang D Y, Chen S. 2024. Labile organic carbon fractions in the rhizosphere contribute to nitrogen and phosphorus uptake in rice under long-term crop rotations and nitrogen application. Applied Soil Ecology, 200, 105459.

Wang M M, Zhang S, Wang G C, Xiao L J, Gu B J, Zheng M H, Niu S L, Yang Y H, Luo Y Q, Zhang G L, Shi Z, Luo Z K. 2025. Increased plant productivity exacerbates subsoil carbon losses under warming via nitrogen mining. Nature Geoscience, 18, 510–517.

Wang X, Sheng L X, Li Y, Jiang H B, Lv Z Y, Qi W, Luo W B. 2022. Soil labile organic carbon indicating seasonal dynamics of soil organic carbon in northeast peatland. Ecological Indicators, 138, 108847.

Wen J L, Li Z Y, Yao J F, Wang S, Liu Y, Liu Y. 2024. The combined application of organic and inorganic fertilizers improved the quality of colored wheat by physicochemical properties and rheological characteristics of starch. International Journal of Biological Macromolecules, 282, 137175.

Wu H Q, Du S Y, Zhang Y L, An J, Zou H T, Zhang Y L, Yu N. 2019. Effects of irrigation and nitrogen fertilization on greenhouse soil organic nitrogen fractions and soil-soluble nitrogen pools. Agricultural Water Management, 216, 415–424.

Wu Y, Zhao D Y, Wang Y H, Wang J P, Wu Y Y, Peng P H, Yang L Y, Wu Y H, Bing H J, Bol R. 2024. Plant species modulate wildfire effects on soil phosphorus fractions in alpine forest of Eastern Tibetan Plateau. Forest Ecology and Management, 573, 122338.

Xia H, Wang J Y, Riaz M, Babar S, Li Y X, Wang X L, Xia X Y, Liu B, Jiang C C. 2024. Co-application of biochar and potassium fertilizer improves soil potassium availability and microbial utilization of organic carbon: A four-year study. Journal of Cleaner Production, 469, 143211.

Xue H P, Zhou X, Chen Y Y, Tu L Y, Liu X Q, Jiang S W, Ji M, Leng C R, Shen Y A. 2025. Effects of historical dust-mediated atmospheric phosphorus deposition on phosphorus fractions in sediments and primary productivity of Dalongwan Maar Lake, Northeast China. Palaeogeography, Palaeoclimatology, Palaeoecology, 660, 112679.

Yang C D, Zhang S H, Tan G Y, Zhu S Q, Wang J. 2024. Effects of cover crops and nitrogen fertilization on soil physical properties, carbon and nitrogen fractions, and winter wheat yield in the Chinese Loess Plateau: A 4-year field experiment. Field Crops Research, 312, 109400.

Yang H K, Xiao Y, Zhang X, Huang X L, Fan G Q. 2022. Maize straw mulching with uniconazole application increases the tillering capacity and grain yield of dryland winter wheat (Triticum aestivum L.). Field Crops Research, 284, 108573.

Yang W J, Yu J, Li Y H, Jia B L, Jiang L G, Yuan A J, Ma Y, Huang M, Cao H B, Liu J S, Qiu W H, Wang Z H. 2024. Optimized NPK fertilizer recommendations based on topsoil available nutrient criteria for wheat in drylands of China. Journal of Integrative Agriculture, 23, 2421–2433.

Yuan S, Li R N, Zhang Y J, Luan H A, Tang J W, Wang L Y, Ji H J, Huang S W. 2024. Effects of long-term partial substitution of inorganic fertilizer with pig manure and/or straw on nitrogen fractions and microbiological properties in greenhouse vegetable soils. Journal of Integrative Agriculture, 23, 2083–2098.

Yuan Z X, Guan Q W, Chen X L, Zou P J, Gu Y Q, Wu Q, Niu Y Y, Meshack A O. 2023. Tree diversity increases soil C and N stocks of secondary forests in subtropical China. Catena, 222, 106812.

Zeng W J, Wang Z D, Chen X Y, Yao X D, Ma Z Q, Wang W. 2023. Nitrogen deficiency accelerates soil organic carbon decomposition in temperate degraded grasslands, Science of the Total Environment, 881, 163424.

Zhai L C, Wang Z B, Zhai Y C, Zhang L H, Zheng M J, Yao H P, Lv L H, Shen H P, Zhang J T, Yao Y R, Jia X L. 2022. Partial substitution of chemical fertilizer by organic fertilizer benefits grain yield, water use efficiency, and economic return of summer maize. Soil & Tillage Research, 217, 105287.

Zhang F, Chen M R, Xing Y Y, Han J, Wang X K. 2025. Reduction of nitrogen fertilizer and simultaneously application of organic fertilizer optimizes yield, water productivity and nitrogen metabolism of spring maize by improving soil properties in the Loess Plateau of China. Journal of Agriculture and Food Research, 19, 101634.

Zhang L H, Li J H, Chen S, Huang W L, Huang L L, Zhu H F, Tang Y X, Li H, You C M, Wang L X, Liu S N, Xu H W, Xu L, Tan B, Xu Z F, Zhu P, Yuan Y L, Yin R, Zhu Y W, Zhang L. 2025. Dynamics of soil phosphorus fractions during the early recovery stage of two typical plantations in China after strip clearcutting. Catena, 250, 108767.

Zhang M Y, Riaz M, Liu B, Xia H., El-desouki Z, Jiang C C. 2020. Two-year study of biochar: Achieving excellent capability of potassium supply via alter clay mineral composition and potassium-dissolving bacteria activity. Science of the Total Environment, 717, 137286.

Zhang Y Z, Resch M C, Schütz M, Liao Z Y, Frey B, Risch A C. 2024. Strengthened plant-microorganism interaction after topsoil removal cause more deterministic microbial assembly processes and increased soil nitrogen mineralization. Science of the Total Environment, 950, 175031.

Zhang Z G, Wang J, Huang W B, Chen J L, Wu F Q, Jia Y Y, Han Y C, Wang G P, Feng L, Li X F, Lei Y P, Yang B F, Xiong S W, Xin M H, Li Y B, Wang Z B. 2022. Cover crops and N fertilization affect soil ammonia volatilization and N2O emission by regulating the soil labile carbon and nitrogen fractions. Agriculture, Ecosystems and Environment, 340, 108188.

Zhang Z T, Luo R Y, Liu Q H, Qiang W, Liang J, Hou E Q, Zhao C Z, Pang X Y. 2023. Linking soil phosphorus fractions to abiotic factors and the microbial community during subalpine secondary succession: Implications for soil phosphorus availability. Catena, 233, 107501. 

Zhao J, Ren T, Fang Y T, Zhang S T, Cong R H, Li X K, Lu Z F, Zhu J, Lu J W. 2025. Soil organic nitrogen sequestrated more in oilseed rape-rice rotation than in wheat-rice rotation under different fertilizations. Agriculture, Ecosystems and Environment, 381, 109445.

Zhao S C, He P, Qiu S J, Jia L L, Liu M C, Jin J Y, Johnston A M. 2014. Long-term effects of potassium fertilization and straw return on soil potassium levels and crop yields in north-central China. Field Crops Research, 169, 116–122.

Zhao Y N, Mao X X, Zhang M S, Yang W, Di H J, Ma L, Liu W J, Li B W. 2021. The application of Bacillus Megaterium alters soil microbial community composition, bioavailability of soil phosphorus and potassium, and cucumber growth in the plastic shed system of North China. Agriculture, Ecosystems and Environment, 307, 107236.

Zhong Y Q W, Yan W M, Shangguan Z P. 2015. Soil carbon and nitrogen fractions in the soil profile and their response to long-term nitrogen fertilization in a wheat field. Catena, 135, 38–46.

Zhou J Y, Gu B J, Schlesinger W H, Ju X T. 2016. Significant accumulation of nitrate in Chinese

Semi-humid croplands. Scientific Reports, 6, 25088.

Zhou Q W, Yan W M, Shanguan Z P. 2015. Soil carbon and nitrogen fractions in the soil profile and their response to long-term nitrogen fertilization in a wheat field. Catena, 135, 38–46.

Zhu J, Li M, Whelan M. 2018. Phosphorus activators contribute to legacy phosphorus availability in agricultural soils: A review. Science of the Total Environment, 612, 522–537.

Zuo H L, Xu W S, Liu Z Y, Smaill S J, Zhou X Q. 2023. Long-term plant diversity increases soil extractable organic carbon and nitrogen contents in a subtropical forest. Science of the Total Environment, 878, 163118.