Akdogan G, Tufekci E D, Uranbey S, Unver T. 2016. miRNA-based drought regulation in wheat. Functional & Integrative Genomics, 16, 221–233.

Albertos P, Tatematsu K, Mateos I, Sánchez-Vicente I, Fernández-Arbaizar A, Nakabayashi K, Nambara E, Godoy M, Franco J M, Solano R, Gerna D, Roach T, Stöggl W, Kranner I, Perea-Resa C, Salinas J, Lorenzo O. 2021. Redox feedback regulation of ANAC089 signaling alters seed germination and stress response. Cell Reports, 35, 109263.

Alshareef N O, Otterbach S L, Allu A D, Woo Y H, De Werk T, Kamranfar I, Mueller-Roeber B, Tester M, Balazadeh S, Schmöckel S M. 2022. NAC transcription factors ATAF1 and ANAC055 affect the heat stress response in Arabidopsis. Scientific Reports, 12, 11264.

An J P, Li R, Qu F J, You C X, Wang X F, Hao Y J. 2018. An apple NAC transcription factor negatively regulates cold tolerance via CBF-dependent pathway. Journal of Plant Physiology, 221, 74–80.

Bian Z, Gao H, Wang C. 2020. NAC transcription factors as positive or negative regulators during ongoing battle between pathogens and our food crops. International Journal of Molecular Sciences, 22, 81.

Cao H, Wang L, Nawaz M A, Niu M, Sun J, Xie J, Kong Q, Huang Y, Cheng F, Bie Z. 2017. Ectopic expression of Pumpkin NAC transcription factor CmNAC1 improves multiple abiotic stress tolerance in Arabidopsis. Frontiers in Plant Science, 8, 2052.

Chang N, Zhou Z, Li Y, Zhang X. 2022. Exogenously applied Spd and Spm enhance drought tolerance in tea plants by increasing fatty acid desaturation and plasma membrane H+-ATPase activity. Plant Physiology and Biochemistry, 170, 225–233.

Chen Q J, Zhang L P, Song S R, Wang L, Xu W P, Zhang C X, Wang S P, Liu H F, Ma C. 2022. vvi-miPEP172b and vvi-miPEP3635b increase cold tolerance of grapevine by regulating the corresponding MIRNA genes. Plant Science, 325, 111450.

Chen Y, Yu X. 2023. Endoplasmic reticulum stress-responsive microRNAs are involved in the regulation of abiotic stresses in wheat. Plant Cell Reports, 42, 1433–1452.

Chi Q, Du L Y, Ma W, Niu R Y, Wu B W, Guo L J, Ma M, Liu X L, Zhao H X. 2022. The miR164-TaNAC14 module regulates root development and abiotic-stress tolerance of wheat seedlings. Journal of Integrative Agriculture, 22, 981–998.

Diao P, Chen C, Zhang Y, Meng Q, Lv W, Ma N. 2020. The role of NAC transcription factor in plant cold response. Plant Signaling & Behavior, 15, 1785668.

Ding D, Zhang L, Wang H, Liu Z, Zhang Z, Zheng Y. 2009. Differential expression of miRNAs in response to salt stress in maize roots. Annals of Botany, 103, 29–38.

Dong Y, Tang M, Huang Z, Song J, Xu J, Ahammed G J, Yu J, Zhou Y. 2022. The miR164a-NAM3 module confers cold tolerance by inducing ethylene production in tomato. Plant Journal, 111, 440–456.

Fang Y, Xie K, Xiong L. 2014. Conserved miR164-targeted NAC genes negatively regulate drought resistance in rice. Journal of Experimental Botany, 65, 2119–2135.

Feng C, Zhang X, Du B, Xiao Y, Wang Y, Sun Y, Zhou X, Wang C, Liu Y, Li T H. 2023. MicroRNA156ab regulates apple plant growth and drought tolerance by targeting transcription factor MsSPL13. Plant Physiology, 192, 1836–1857.

Guo Z, Kuang Z, Wang Y, Zhao Y, Tao Y, Cheng C, Yang J, Lu X, Hao C, Wang T, Cao X, Wei J, Li L, Yang X. 2020. PmiREN: A comprehensive encyclopedia of plant miRNAs. Nucleic Acids Research, 48, 1114–1121.

Hao Y J, Wei W, Song Q X, Chen H W, Zhang Y Q, Wang F, Zou H F, Lei G, Tian A G, Zhang W K, Ma B, Zhang J S, Chen S Y. 2011. Soybean NAC transcription factors promote abiotic stress tolerance and lateral root formation in transgenic plants. Plant Journal, 68, 302–313.

Han K, Zhao Y, Sun Y, Li Y. 2023. NACs, generalist in plant life. Plant Biotechnology Journal, 21, 2433–2457.

Hernandez Y, Goswami K, Sanan-Mishra N. 2020. Stress induced dynamic adjustment of conserved miR164:NAC module. Plant Environment Interactions, 1, 134–151.

Hou X M, Zhang H F, Liu S Y, Wang X K, Zhang Y M, Meng Y C, Luo D, Chen R G. 2020. The NAC transcription factor CaNAC064 is a regulator of cold stress tolerance in peppers. Plant Science, 291, 110346.

Hu P, Zhang K, Yang C. 2019. BpNAC012 positively regulates abiotic stress responses and secondary wall biosynthesis. Plant Physiology, 179, 700–717.

Huang J C, Piater L A, Dubery I A. 2012. The NAC transcription factor gene ANAC072 is differentially expressed in Arabidopsis thaliana in response to microbe-associated molecular pattern (MAMP) molecules. Physiological and Molecular Plant Pathology, 80, 19–27.

Huang S Q, Xiang A L, Che L L, Chen S, Li H, Song J B, Yang Z M. 2010. A set of miRNAs from Brassica napus in response to sulphate deficiency and cadmium stress. Plant Biotechnology Journal, 8, 887–899.

Jeyaraj A, Wang X, Wang S, Liu S, Zhang R, Wu A, Wei C. 2019. Identification of regulatory networks of microRNAs and their targets in response to colletotrichum gloeosporioides in tea plant (Camellia sinensis L.). Frontiers in Plant Science, 10, 1096.

Jia X, Zeng Z, Lyu Y, Zhao S. 2022. Drought-responsive NAC transcription factor RcNAC72 is recognized by RcABF4, interacts with RcDREB2A to enhance drought tolerance in Arabidopsis. International Journal of Molecular Sciences, 23, 1755.

Jin C, Li K Q, Xu X Y, Zhang H P, Chen H X, Chen Y H, Hao J, Wang Y, Huang X S, Zhang S L. 2017. A novel NAC transcription factor, PbeNAC1, of Pyrus betulifolia confers cold and drought tolerance via interacting with PbeDREBs and activating the expression of stress-responsive genes. Frontiers in Plant Science, 8, 1049.

Ju Y L, Yue X F, Min Z, Wang X H, Fang Y L, Zhang J X. 2020. VvNAC17, a novel stress-responsive grapevine (Vitis vinifera L.) NAC transcription factor, increases sensitivity to abscisic acid and enhances salinity, freezing, and drought tolerance in transgenic Arabidopsis. Plant Physiology and Biochemistry, 146, 98–111.

Kong L, Zhu X X, Wang Y W, Xie X F, Jiang C J, Li Y Y. 2018. Identification and expression analysis of tea plant (Camellia sinesis) miR164a and its target gene. Journal of Tea Science, 38, 547–558. (in Chinese)

Kong X M, Zhou Q, Zhou X, Wei B D, Ji S J. 2020. Transcription factor CaNAC1 regulates low-temperature-induced phospholipid degradation in green bell pepper. Journal of Experimental Botany, 71, 1078–1091.

Lee M H, Jeon H S, Kim H G, Park O K. 2017. An Arabidopsis NAC transcription factor NAC4 promotes pathogen-induced cell death under negative regulation by microRNA164. New Phytologist, 214, 343–360.

Li J, Lai T, Song H, Xu X. 2017. MiR164 is involved in delaying senescence of strawberry (Fragaria ananassa) fruit by negatively regulating NAC transcription factor genes under low temperature. Russian Journal of Plant Physiology, 64, 251–259.

Li P, Fu J, Xu Y, Shen Y, Zhang Y, Ye Z, Tong W, Zeng X, Yang J, Tang D, Li P, Zuo H, Wu Q, Xia E, Wang S, Zhao J. 2022. CsMYB1 integrates the regulation of trichome development and catechins biosynthesis in tea plant domestication. New Phytologist, 234, 902–917.

Liu G F, Liu J J, He Z R, Wang F M, Yang H, Yan Y F, Gao M J, Gruber M Y, Wan X C, Wei S. 2018. Implementation of CsLIS/NES in linalool biosynthesis involves transcript splicing regulation in Camellia sinensis. Plant Cell & Environment, 41, 176–186.

Liu L, Chen H, Zhu J, Tao L, Wei C. 2022. miR319a targeting of CsTCP10 plays an important role in defense against gray blight disease in tea plant (Camellia sinensis). Tree Physiology, 42, 1450–1462.

Liu Y, Sun J, Wu Y. 2016. Arabidopsis ATAF1 enhances the tolerance to salt stress and ABA in transgenic rice. Journal of Plant Research, 129, 955–962.

Luo J, Huang S, Chang Y, Li H, Guo G. 2023. Physiological and transcriptomic analyses reveal tea plant (Camellia sinensis L.) adapts to extreme freezing stress during winter by regulating cell wall structure. BMC Genomics, 24, 558.

Ma L, Zhao Y, Chen M, Li Y, Shen Z, Cao Y, Wu D, Yu M, Grierson D, Shi Y, Chen K. 2023. The microRNA ppe-miR393 mediates auxin-induced peach fruit softening by promoting ethylene production. Plant Physiology, 192, 1638–1655.

Ma W, Kang X, Liu P, She K, Zhang Y, Lin X, Li B, Chen Z. 2022. The NAC-like transcription factor CsNAC7 positively regulates the caffeine biosynthesis-related gene yhNMT1 in Camellia sinensis. Horticulture Research, 9, 46.

Mao H, Li S, Wang Z, Cheng X, Li F, Mei F, Chen N, Kang Z. 2020. Regulatory changes in TaSNAC8-6A are associated with drought tolerance in wheat seedlings. Plant Biotechnology Journal, 18, 1078–1092.

Mei C, Yang J, Mei Q, Jia D, Yan P, Feng B, Mamat A, Gong X, Guan Q, Mao K, Wang J, Ma F. 2023. MdNAC104 positively regulates apple cold tolerance via CBF-dependent and CBF-independent pathways. Plant Biotechnology Journal, 21, 2057–2073.

Otte K A, Nolte V, Mallard F, Schlötterer C. 2021. The genetic architecture of temperature adaptation is shaped by population ancestry and not by selection regime. Genome Biology, 22, 211.

Peng H, Phung J, Stowe E C, Dhingra A, Neff M M. 2022. The NAC transcription factor ATAF2 promotes ethylene biosynthesis and response in Arabidopsis thaliana seedlings. FEBS Letters, 596, 1586–1599.

Peng J, Li N, Di T, Ding C, Li X, Wu Y, Hao X, Wang Y, Yang Y, Wang X, Wang L. 2022. The interaction of CsWRKY4 and CsOCP3 with CsICE1 regulates CsCBF1/3 and mediates stress response in tea plant (Camellia sinensis). Environmental and Experimental Botany, 199, 104892.

Pi X T, Chang N, Zhou Z W, Li Y Y, Zhang X C. 2023. CsFAD2 and CsFAD5 are key genes for C18:2 fatty acid pathway-mediated cold tolerance in tea (Camellia sinensis). Environmental and Experimental Botany, 210, 105317.

Shah S T, Pang C, Fan S, Song M, Arain S, Yu S. 2013. Isolation and expression profiling of GhNAC transcription factor genes in cotton (Gossypium hirsutum L.) during leaf senescence and in response to stresses. Gene, 531, 220–234.

Shan W, Kuang J F, Lu W J, Chen J Y. 2014. Banana fruit NAC transcription factor MaNAC1 is a direct target of MaICE1 and involved in cold stress through interacting with MaCBF1. Plant Cell & Environment, 37, 2116–2127.

Sharma R, Upadhyay S, Bhattacharya S, Singh A. 2021. Abiotic stress-responsive miRNA and transcription factor-mediated gene regulatory network in Oryza sativa: Construction and structural measure study. Frontiers in Genetics, 12, 618089.

Shen X, He J, Ping Y, Guo J, Hou N, Cao F, Li X, Geng D, Wang S, Chen P, Qin G, Ma F, Guan Q. 2022. The positive feedback regulatory loop of miR160-Auxin Response Factor 17-HYPONASTIC LEAVES 1 mediates drought tolerance in apple trees. Plant Physiology, 188, 1686–1708.

Shim J S, Oh N, Chung P J, Kim Y S, Choi Y D, Kim J K. 2018. Overexpression of OsNAC14 improves drought tolerance in rice. Frontiers in Plant Science, 9, 310.

Singh P, Dutta P, Chakrabarty D. 2021. miRNAs play critical roles in response to abiotic stress by modulating cross-talk of phytohormone signaling. Plant Cell Reports, 40, 1617–1630.

Song S S, Ran W X, Gao L H, Wang Y C, Lv W Y, Tao Y, Chen L, Li C F. 2024. A functional study reveals CsNAC086 regulated the biosynthesis of flavonols in Camellia sinensis. Planta, 259, 147.

Strzyz P. 2021. microRNA communication in plants. Nature Reviews Molecular Cell Biology, 22, 775.

Sukiran N L, Ma J C, Ma H, Su Z. 2019. ANAC019 is required for recovery of reproductive development under drought stress in Arabidopsis. Plant Molecular Biology, 99, 161–174.

Tang Y, Lu L, Huang X, Zhao D, Tao J. 2023. The herbaceous peony transcription factor WRKY41a promotes secondary cell wall thickening to enhance stem strength. Plant Physiology, 191, 428–445.

Wan X, Wang Z, Duan W, Huang T, Song H, Xu X. 2023. Knockdown of Sly-miR164a enhanced plant salt tolerance and improved preharvest and postharvest fruit nutrition of tomato. International Journal of Molecular Sciences, 24, 4639.

Wang L, Zhao H, Chen D, Li L, Sun H, Lou Y, Gao Z. 2016. Characterization and primary functional analysis of a bamboo NAC gene targeted by miR164b. Plant Cell Reports, 35, 1371–1383.

Wang M, Ren L T, Wei X Y, Ling Y M, Gu H T, Wang S S, Ma X F, Kong G C. 2022. NAC transcription factor TwNAC01 positively regulates drought stress responses in Arabidopsis and Triticale. Frontiers in Plant Science, 13, 877016.

Wang P, Yang Y, Shi H, Wang Y, Ren F. 2019. Small RNA and degradome deep sequencing reveal respective roles of cold-related microRNAs across Chinese wild grapevine and cultivated grapevine. BMC Genomics, 20, 740.

Wang Q, Hu F, Yao Z, Zhao X, Chu G, Ye J. 2023. Comprehensive genomic characterisation of the NAC transcription factor family and its response to drought stress in Eucommia ulmoides. PeerJ, 11, e16298.

Wang Y, Jiang C J, Li Y Y, Wei C L, Deng W W. 2012. CsICE1 and CsCBF1: Two transcription factors involved in cold responses in Camellia sinensis. Plant Cell Reports, 31, 27–34.

Wang Y, Samarina L, Mallano A I, Tong W, Xia E. 2023. Recent progress and perspectives on physiological and molecular mechanisms underlying cold tolerance of tea plants. Frontiers in Plant Science, 14, 1145609.

Wen L, Liu T, Deng Z, Zhang Z, Wang Q, Wang W, Li W, Guo Y. 2022. Characterization of NAC transcription factor NtNAC028 as a regulator of leaf senescence and stress responses. Frontiers in Plant Science, 13, 941026.

Xia L, Sun S, Han B, Yang X. 2023. NAC domain transcription factor gene GhNAC3 confers drought tolerance in plants. Plant Physiology and Biochemistry, 195, 114–123.

Xie J, Chen Y, Cai G, Cai R, Hu Z, Wang H. 2023. Tree visualization by one table (tvBOT): A web application for visualizing, modifying and annotating phylogenetic trees. Nucleic Acids Research, 51, W587–W592.

Yang X, Kim M Y, Ha J, Lee S H. 2019. Overexpression of the soybean NAC gene GmNAC109 increases lateral Root formation and abiotic stress tolerance in transgenic Arabidopsis plants. Frontiers in Plant Science, 10, 1036.

Yong Y, Zhang Y, Lyu Y. 2019. A stress-responsive NAC transcription factor from tiger lily (LlNAC2) interacts with LlDREB1 and LlZHFD4 and enhances various abiotic stress tolerance in Arabidopsis. International Journal of Molecular Sciences, 20, 3225.

Zhang X, Cao X, Xia Y, Ban Q, Cao L, Li S, Li Y. 2022a. CsCBF5 depletion impairs cold tolerance in tea plants. Plant Science, 325, 111463.

Zhang X, Li L, Lang Z, Li D, He Y, Zhao Y, Tao H, Wei J, Li Q, Hong G. 2022b. Genome-wide characterization of NAC transcription factors in Camellia sinensis and the involvement of CsNAC28 in drought tolerance. Frontiers in Plant Science, 13, 1065261.

Zhang X, Wang W, Wang M, Zhang H Y, Liu J H. 2016. The miR396b of Poncirus trifoliata functions in cold tolerance by regulating ACC oxidase gene expression and modulating ethylene-polyamine homeostasis. Plant & Cell Physiology, 57, 1865–1878.

Zhang X, Xia Y, Li S, Cao L, Zhou Z, Chang N, Li Y. 2023. The arginine decarboxylase gene CsADC1, associated with the polyamine pathway, plays an important role in tea cold tolerance. Environmental and Experimental Botany, 214, 105473.

Zhang Y, Zhu X, Chen X, Song C, Zou Z, Wang Y, Wang M, Fang W, Li X. 2014. Identification and characterization of cold-responsive microRNAs in tea plant (Camellia sinensis) and their targets using high-throughput sequencing and degradome analysis. BMC Plant Biology, 14, 271.

Zhao M, Jin J, Wang J, Gao T, Luo Y, Jing T, Hu Y, Pan Y, Lu M, Schwab W, Song C. 2022. Eugenol functions as a signal mediating cold and drought tolerance via UGT71A59-mediated glucosylation in tea plants. Plant Journal, 109, 1489–1506.

Zhao M, Wang L, Wang J, Jin J, Zhang N, Lei L, Gao T, Jing T, Zhang S, Wu Y, Wu B, Hu Y, Wan X, Schwab W, Song C. 2020. Induction of priming by cold stress via inducible volatile cues in neighboring tea plants. Journal of Integrative Plant Biology, 62, 1461–1468.

Zhao X, Li P, Zuo H, Peng A, Lin J, Li P, Wang K, Tang Q, Tadege M, Liu Z, Zhao J. 2023. CsMYBL2 homologs modulate the light and temperature stress-regulated anthocyanin and catechins biosynthesis in tea plants (Camellia sinensis). Plant Journal, 115, 1051–1070.

Zheng G, Wei W, Li Y, Kan L, Wang F, Zhang X, Li F, Liu Z, Kang C. 2019. Conserved and novel roles of miR164-CUC2 regulatory module in specifying leaf and floral organ morphology in strawberry. New Phytologist, 224, 480–492.