Antuña E, Cachán-Vega C, Bermejo-Millo J C, Potes Y, Caballero B, Vega-Naredo I, Coto-Montes A, Garcia-Gonzalez C. 2022. Inflammaging: Implications in sarcopenia. International Journal of Molecular Sciences, 23, 15039.

Baccarin R Y A, Seidel S R T, Michelacci Y M, Tokawa P K A, Oliveira T M. 2022. Osteoarthritis: A common disease that should be avoided in the athletic horse's life. Animal Frontiers : the Review Magazine of Animal Agriculture, 12, 25-36.

Basse A L, Agerholm M, Farup J, Dalbram E, Nielsen J, Ørtenblad N, Altıntaş A, Ehrlich A M, Krag T, Bruzzone S, Dall M, De Guia R M, Jensen J B, Møller A B, Karlsen A, Kjær M, Barrès R, Vissing J, Larsen S, Jessen N, et al. 2021. Nampt controls skeletal muscle development by maintaining Ca²⁺ homeostasis and mitochondrial integrity. Molecular Metabolism, 53, 101271.

Bou T, Ding W, Ren X, Liu H, Gong W, Jia Z, Zhang X, Dugarjaviin M, Bai D. 2024. Muscle fibre transition and transcriptional changes of horse skeletal muscles during traditional Mongolian endurance training. Equine Veterinary Journal, 56, 178-192.

Bu D, Luo H, Huo P, Wang Z, Zhang S, He Z, Wu Y, Zhao L, Liu J, Guo J, Fang S, Cao W, Yi L, Zhao Y, Kong L. 2021. KOBAS-i: Intelligent prioritization and exploratory visualization of biological functions for gene enrichment analysis. Nucleic Acids Research, 49, W317-W325.

Cai C, Wan P, Wang H, Cai X, Wang J, Chai Z, Wang J, Wang H, Zhang M, Yang N, Wu Z, Zhu J, Yang X, Li Y, Yue B, Dang R, Zhong J. 2023. Transcriptional and open chromatin analysis of bovine skeletal muscle development by single-cell sequencing. Cell Proliferation, 56, e13430.

Camillo F, Rota A, Biagini L, Tesi M, Fanelli D, Panzani D. 2018. The current situation and trend of donkey industry in Europe. Journal of Equine Veterinary Science, 65, 44-49.

Chen C, Sun Y, Liang H, Yu D, Hu S. 2017. A meta-analysis of the association of CKM gene rs8111989 polymorphism with sport performance. Biology of Sport, 34, 323-330.

Chen L, Bai J, Li Y. 2020. miR‑29 mediates exercise‑induced skeletal muscle angiogenesis by targeting VEGFA, COL4A1 and COL4A2 via the PI3K/Akt signaling pathway. Molecular Medicine Reports, 22, 661-670.

Chen X, Liang D, Huang Z, Jia G, Zhao H, Liu G. 2021. Quercetin regulates skeletal muscle fiber type switching via adiponectin signaling. Food & Function, 12, 2693-2702.

Chen X, Wu H, Liu Y, Liu L, Houser S R, Wang W E. 2024. Metabolic reprogramming: A byproduct or a driver of cardiomyocyte proliferation? Circulation, 149, 1598-1610.

Dietrich J, Handschuh S, Steidl R, Böhler A, Forstenpointner G, Egerbacher M, Peham C, Schöpper H. 2021. muscle fibre architecture of thoracic and lumbar longissimus dorsi muscle in the horse. Animals : an Open Access Journal From MDPI, 11, 915.

Fastman N M, Liu Y, Ramanan V, Merritt H, Ambing E, Depaoli-Roach A A, Roach P J, Hurley T D, Mellem K T, Ullman J C, Green E, Morgans D, Tzitzilonis C. 2022. The structural mechanism of human glycogen synthesis by the GYS1-GYG1 complex. Cell Reports, 40, 111041.

Felix M R, Turner R M, Dobbie T, Hinrichs K. 2022. Successful in vitro fertilization in the horse: Production of blastocysts and birth of foals after prolonged sperm incubation for capacitation. Biology of Reproduction, 107, 1551-1564.

Fritz P, Fritz R, Bóday P, Bóday Á, Bató E, Kesserű P, Oláh C. 2024. Gut microbiome composition: link between sports performance and protein absorption? Journal of the International Society of Sports Nutrition, 21, 2297992.

Fujimaki S, Matsumoto T, Muramatsu M, Nagahisa H, Horii N, Seko D, Masuda S, Wang X, Asakura Y, Takahashi Y, Miyamoto Y, Usuki S, Yasunaga K-I, Kamei Y, Nishinakamura R, Minami T, Fukuda T, Asakura A, Ono Y. 2022. The endothelial Dll4-muscular Notch2 axis regulates skeletal muscle mass. Nature Metabolism, 4, 180-189.

Giarratana N, Conti F, La Rovere R, Gijsbers R, Carai P, Duelen R, Vervliet T, Bultynck G, Ronzoni F, Piciotti R, Costamagna D, Fulle S, Barravecchia I, Angeloni D, Torrente Y, Sampaolesi M. 2020. MICAL2 is essential for myogenic lineage commitment. Cell Death & Disease, 11, 654.

Giuliani G, Rosina M, Reggio A. 2022. Signaling pathways regulating the fate of fibro/adipogenic progenitors (FAPs) in skeletal muscle regeneration and disease. The FEBS Journal, 289, 6484-6517.

Gu S, Huang Q, Jie Y, Sun C, Wen C, Yang N. 2024. Transcriptomic and epigenomic landscapes of muscle growth during the postnatal period of broilers. Journal of Animal Science and Biotechnology, 15, 91.

Guo L, Han M, Xu J, Zhou W, Shi H, Chen S, Pang W, Zhang X, Duan Y, Yin Y, Li F. 2025. snRNA-seq and spatial transcriptome reveal cell-cell crosstalk mediated metabolic regulation in porcine skeletal muscle. Journal of Cachexia, Sarcopenia and Muscle, 16, e13752.

Hitsumoto T, Tsukamoto O, Matsuoka K, Li J, Liu L, Kuramoto Y, Higo S, Ogawa S, Fujino N, Yoshida S, Kioka H, Kato H, Hakui H, Saito Y, Okamoto C, Inoue H, Hyejin J, Ueda K, Segawa T, Nishimura S, et al. 2023. Restoration of cardiac myosin light chain kinase ameliorates systolic dysfunction by reducing superrelaxed myosin. Circulation, 147, 1902-1918.

Honda M, Inoue R, Nishiyama K, Ueda T, Komuro A, Amano H, Sugisawa R, Dash S, Shirakawa J, Okada H. 2024. Vgll2 as an integrative regulator of mitochondrial function and contractility specific to skeletal muscle. Journal of Cellular Physiology, 239, e31436.

Iwabu M, Yamauchi T, Okada-Iwabu M, Sato K, Nakagawa T, Funata M, Yamaguchi M, Namiki S, Nakayama R, Tabata M, Ogata H, Kubota N, Takamoto I, Hayashi Y K, Yamauchi N, Waki H, Fukayama M, Nishino I, Tokuyama K, Ueki K, et al. 2010. Adiponectin and AdipoR1 regulate PGC-1alpha and mitochondria by Ca²⁺ and AMPK/SIRT1. Nature, 464, 1313-1319.

Jia M, Yue X, Sun W, Zhou Q, Chang C, Gong W, Feng J, Li X, Zhan R, Mo K, Zhang L, Qian Y, Sun Y, Wang A, Zou Y, Chen W, Li Y, Huang L, Yang Y, Zhao Y, et al. 2023. ULK1-mediated metabolic reprogramming regulates Vps34 lipid kinase activity by its lactylation. Science Advances, 9, eadg4993.

Jin S, Guerrero-Juarez C F, Zhang L, Chang I, Ramos R, Kuan C-H, Myung P, Plikus M V, Nie Q. 2021. Inference and analysis of cell-cell communication using CellChat. Nature Communications, 12, 1088.

Jin S, Kim J, Willert T, Klein-Rodewald T, Garcia-Dominguez M, Mosqueira M, Fink R, Esposito I, Hofbauer L C, Charnay P, Kieslinger M. 2014. Ebf factors and MyoD cooperate to regulate muscle relaxation via Atp2a1. Nature Communications, 5, 3793.

Kanehisa M, Goto S. 2000. KEGG: kyoto encyclopedia of genes and genomes. Nucleic Acids Research, 28, 27-30.

Kedlian V R, Wang Y, Liu T, Chen X, Bolt L, Tudor C, Shen Z, Fasouli E S, Prigmore E, Kleshchevnikov V, Pett J P, Li T, Lawrence J E G, Perera S, Prete M, Huang N, Guo Q, Zeng X, Yang L, Polański K, et al. 2024. Human skeletal muscle aging atlas. Nature Aging, 4, 727-744.

Korsunsky I, Millard N, Fan J, Slowikowski K, Zhang F, Wei K, Baglaenko Y, Brenner M, Loh P-R, Raychaudhuri S. 2019. Fast, sensitive and accurate integration of single-cell data with Harmony. Nature Methods, 16, 1289-1296.

Lai Y, Ramírez-Pardo I, Isern J, An J, Perdiguero E, Serrano A L, Li J, García-Domínguez E, Segalés J, Guo P, Lukesova V, Andrés E, Zuo J, Yuan Y, Liu C, Viña J, Doménech-Fernández J, Gómez-Cabrera M C, Song Y, Liu L, et al. 2024. Multimodal cell atlas of the ageing human skeletal muscle. Nature, 629, 154-164.

Li C, Li X, Guo R, Ni W, Liu K, Liu Z, Dai J, Xu Y, Abduriyim S, Wu Z, Zeng Y, Lei B, Zhang Y, Wang Y, Zeng W, Zhang Q, Chen C, Qiao J, Liu C, Hu S. 2023. Expanded catalogue of metagenome-assembled genomes reveals resistome characteristics and athletic performance-associated microbes in horse. Microbiome, 11, 7.

Li X, Ma J, Li H, Li H, Ma Y, Deng H, Yang K. 2024. Effect of β-alanine on the athletic performance and blood amino acid metabolism of speed-racing Yili horses. Frontiers In Veterinary Science, 11, 1339940.

Li Y, Pan H, Zhang X, Wang H, Liu S, Zhang H, Qian H, Wang L, Ying H. 2019. Geniposide improves glucose homeostasis via regulating FoxO1/PDK4 in skeletal muscle. Journal of Agricultural and Food Chemistry, 67, 4483-4492.

Lin Z, Tian H, Lam K S L, Lin S, Hoo R C L, Konishi M, Itoh N, Wang Y, Bornstein S R, Xu A, Li X. 2013. Adiponectin mediates the metabolic effects of FGF21 on glucose homeostasis and insulin sensitivity in mice. Cell Metabolism, 17, 779-789.

Liu B, Li C, Li Z, Wang D, Ren X, Zhang Z. 2020. An entropy-based metric for assessing the purity of single cell populations. Nature Communications, 11, 3155.

Liu X, Zhang Y, Liu W, Li Y, Pan J, Pu Y, Han J, Orlando L, Ma Y, Jiang L. 2022. A single-nucleotide mutation within the TBX3 enhancer increased body size in Chinese horses. Current Biology : CB, 32, 480-487.

Liu Y, Wu X, Xu Q, Lan X, Li W. 2024. Temporal transcriptome dynamics of longissimus dorsi reveals the mechanism of the differences in muscle development and IMF deposition between fuqing goats and Nubian goats. Animals : An Open Access Journal From MDPI, 14, 1770.

Love M I, Huber W, Anders S. 2014. Moderated estimation of fold change and dispersion for RNA-seq data with DESeq2. Genome Biology, 15, 550.

Lu W, Feng W, Lai J, Yuan D, Xiao W, Li Y. 2023. Role of adipokines in sarcopenia. Chinese Medical Journal, 136, 1794-1804.

Massengill M T, Ashraf H M, Chowdhury R R, Chrzanowski S M, Kar J, Warren S A, Walter G A, Zeng H, Kang B-H, Anderson R H, Moss R L, Kasahara H. 2016. Acute heart failure with cardiomyocyte atrophy induced in adult mice by ablation of cardiac myosin light chain kinase. Cardiovascular Research, 111, 34-43.

Matarneh S K, Silva S L, Gerrard D E. 2021. New insights in muscle biology that alter meat quality. Annual Review of Animal Biosciences, 9, 355-377.

Mickelson J R, Valberg S J. 2015. The genetics of skeletal muscle disorders in horses. Annual Review of Animal Biosciences, 3, 197-217.

Murgia M, Nogara L, Baraldo M, Reggiani C, Mann M, Schiaffino S. 2021. Protein profile of fiber types in human skeletal muscle: A single-fiber proteomics study. Skeletal Muscle, 11, 24.

Nolte W, Thaller G, Kuehn C. 2019. Selection signatures in four German warmblood horse breeds: Tracing breeding history in the modern sport horse. PLoS ONE, 14, e0215913.

Otten B M J, Sthijns M M J P E, Troost F J. 2023. A combination of acetate, propionate, and butyrate increases glucose uptake in C2C12 myotubes. Nutrients, 15, 946.

Pan X, Li X, Dong L, Liu T, Zhang M, Zhang L, Zhang X, Huang L, Shi W, Sun H, Fang Z, Sun J, Huang Y, Shao H, Wang Y, Yin M. 2024. Tumour vasculature at single-cell resolution. Nature, 632, 429-436.

Petrany M J, Swoboda C O, Sun C, Chetal K, Chen X, Weirauch M T, Salomonis N, Millay D P. 2020. Single-nucleus RNA-seq identifies transcriptional heterogeneity in multinucleated skeletal myofibers. Nature Communications, 11, 6374.

Poškienė I, Gruodytė R, Autukaitė J, Juozaitienė V, Antanaitis R. 2021. Speed and blood parameters differ between Arabian and Žemaitukai horses during endurance racing. Animals: an Open Access Journal From MDPI, 11, 995.

Reich P, Möller S, Stock K F, Nolte W, Von Depka Prondzinski M, Reents R, Kalm E, Kühn C, Thaller G, Falker-Gieske C, Tetens J. 2024. Genomic analyses of withers height and linear conformation traits in German Warmblood horses using imputed sequence-level genotypes. Genetics, Selection, Evolution : GSE, 56, 45.

Ribbenstedt A, Ziarrusta H, Benskin J P. 2018. Development, characterization and comparisons of targeted and non-targeted metabolomics methods. PLoS ONE, 13, e0207082.

Risebro C A, Searles R G, Melville A a D, Ehler E, Jina N, Shah S, Pallas J, Hubank M, Dillard M, Harvey N L, Schwartz R J, Chien K R, Oliver G, Riley P R. 2009. Prox1 maintains muscle structure and growth in the developing heart. Development (Cambridge, England), 136, 495-505.

Sales K M, Reimer R A. 2023. Unlocking a novel determinant of athletic performance: The role of the gut microbiota, short-chain fatty acids, and "biotics" in exercise. Journal of Sport and Health Science, 12, 36-44.

Sartori R, Hagg A, Zampieri S, Armani A, Winbanks C E, Viana L R, Haidar M, Watt K I, Qian H, Pezzini C, Zanganeh P, Turner B J, Larsson A, Zanchettin G, Pierobon E S, Moletta L, Valmasoni M, Ponzoni A, Attar S, Da Dalt G, et al. 2021a. Perturbed BMP signaling and denervation promote muscle wasting in cancer cachexia. Science Translational Medicine, 13, 9592.

Sartori R, Romanello V, Sandri M. 2021b. Mechanisms of muscle atrophy and hypertrophy: implications in health and disease. Nature Communications, 12, 330.

Satija R, Farrell J A, Gennert D, Schier A F, Regev A. 2015. Spatial reconstruction of single-cell gene expression data. Nature Biotechnology, 33, 495-502.

Scheiman J, Luber J M, Chavkin T A, Macdonald T, Tung A, Pham L-D, Wibowo M C, Wurth R C, Punthambaker S, Tierney B T, Yang Z, Hattab M W, Avila-Pacheco J, Clish C B, Lessard S, Church G M, Kostic A D. 2019. Meta-omics analysis of elite athletes identifies a performance-enhancing microbe that functions via lactate metabolism. Nature Medicine, 25, 1104-1109.

Semanchik P L, Wesolowski L T, Ryan P J, White-Springer S H, Fluckey J D. 2023. Influence of mechanistic target of rapamycin (mTOR)-regulated anabolic pathways on equine skeletal muscle health. Journal of Equine Veterinary Science, 124, 104281.

Sevane N, Dunner S, Boado A, Cañon J. 2016. Polymorphisms in ten candidate genes are associated with conformational and locomotive traits in Spanish Purebred horses. Journal of Applied Genetics, 58, 355-361.

Slyper M, Porter C B M, Ashenberg O, Waldman J, Drokhlyansky E, Wakiro I, Smillie C, Smith-Rosario G, Wu J, Dionne D, Vigneau S, Jané-Valbuena J, Tickle T L, Napolitano S, Su M-J, Patel A G, Karlstrom A, Gritsch S, Nomura M, Waghray A, et al. 2020. A single-cell and single-nucleus RNA-Seq toolbox for fresh and frozen human tumors. Nature Medicine, 26, 792-802.

Son J S, Chae S A, Zhao L, Wang H, De Avila J M, Zhu M J, Jiang Z, Du M. 2022. Maternal exercise intergenerationally drives muscle-based thermogenesis via activation of apelin-AMPK signaling. EBioMedicine, 76, 103842.

Song S, Wang S, Li N, Chang S, Dai S, Guo Y, Wu X, Cheng Y, Zeng S. 2023a. Genome-wide association study to identify SNPs and candidate genes associated with body size traits in donkeys. Frontiers In Genetics, 14, 1112377.

Song X, Jiang H, Lv P, Cui K, Liu Q, Yin S, Liu H, Li Z. 2023b. Transcriptome analyses reveal transcriptional profiles of horse oocytes before and after in vitro maturation. Reproduction In Domestic Animals=Zuchthygiene, 58, 1468-1479.

Stefaniuk-Szmukier M, Szmatoła T, Ropka-Molik K. 2025. Molecular signatures of exercise Adaptation in Arabian racing horses: Transcriptomic insights from blood and muscle. Genes, 16, 431.

Tan V P, Miyamoto S. 2015. HK2/hexokinase-II integrates glycolysis and autophagy to confer cellular protection. Autophagy, 11, 963-964.

Trapnell C, Cacchiarelli D, Grimsby J, Pokharel P, Li S, Morse M, Lennon N J, Livak K J, Mikkelsen T S, Rinn J L. 2014. The dynamics and regulators of cell fate decisions are revealed by pseudotemporal ordering of single cells. Nature Biotechnology, 32, 381-386.

Umek N, Meznarič M, Šink Ž, Blagotinšek Cokan K, Prosenc Zmrzljak U, Horvat S. 2025. In situ spatial transcriptomic analysis of human skeletal muscle using the Xenium platform. Cell and Tissue Research, 399, 291-302.

Velie B D, Fegraeus K J, Solé M, Rosengren M K, Røed K H, Ihler C-F, Strand E, Lindgren G. 2018. A genome-wide association study for harness racing success in the Norwegian-Swedish coldblooded trotter reveals genes for learning and energy metabolism. BMC Genetics, 19, 80.

Wang G, Wang F, Pei H, Li M, Bai F, Lei C, Dang R. 2022. Genome-wide analysis reveals selection signatures for body size and drought adaptation in Liangzhou donkey. Genomics, 114, 110476.

Wolfsberger W W, Ayala N M, Castro-Marquez S O, Irizarry-Negron V M, Potapchuk A, Shchubelka K, Potish L, Majeske A J, Oliver L F, Lameiro A D, Martínez-Cruzado J C, Lindgren G, Oleksyk T K. 2022. Genetic diversity and selection in Puerto Rican horses. Scientific Reports, 12, 515.

Wouters C P, Toquet M-P, Renaud B, François A-C, Fortier-Guillaume J, Marcillaud-Pitel C, Boemer F, De Tullio P, Richard E A, Votion D-M. 2021. Metabolomic signatures discriminate horses with clinical signs of atypical myopathy from healthy co-grazing horses. Journal of Proteome Research, 20, 4681-4692.

Xia W, Wan Y, Li Y-Y, Zeng H, Lv Z, Li G, Wei Z, Xu S-Q. 2011. PFOS prenatal exposure induce mitochondrial injury and gene expression change in hearts of weaned SD rats. Toxicology, 282, 23-29.

Xiang C, Zhang Y, Chen Q, Sun A, Peng Y, Zhang G, Zhou D, Xie Y, Hou X, Zheng F, Wang F, Gan Z, Chen S, Liu G. 2021. Increased glycolysis in skeletal muscle coordinates with adipose tissue in systemic metabolic homeostasis. Journal of Cellular and Molecular Medicine, 25, 7840-7854.

Xiong Y, Miyamoto N, Shibata K, Valasek M A, Motoike T, Kedzierski R M, Yanagisawa M. 2004. Short-chain fatty acids stimulate leptin production in adipocytes through the G protein-coupled receptor GPR41. Proceedings of the National Academy of Sciences of the United States of America, 101, 1045-1050.

Xu D, Wan B, Qiu K, Wang Y, Zhang X, Jiao N, Yan E, Wu J, Yu R, Gao S, Du M, Liu C, Li M, Fan G, Yin J. 2023. Single-Cell RNA-sequencing provides insight into skeletal muscle evolution during the selection of muscle characteristics. Advanced Science (Weinheim, Baden-Wurttemberg, Germany), e2305080.

Yoshida T, Delafontaine P. 2020. Mechanisms of IGF-1-Mediated Regulation of Skeletal Muscle Hypertrophy and atrophy. Cells, 9, 1970.

Zheng G X Y, Terry J M, Belgrader P, Ryvkin P, Bent Z W, Wilson R, Ziraldo S B, Wheeler T D, Mcdermott G P, Zhu J, Gregory M T, Shuga J, Montesclaros L, Underwood J G, Masquelier D A, Nishimura S Y, Schnall-Levin M, Wyatt P W, Hindson C M, Bharadwaj R, et al. 2017. Massively parallel digital transcriptional profiling of single cells. Nature Communications, 8, 14049.