Ali H E, Reineking B, Münkemüller T. 2017. Effects of plant functional traits on soil stability: Intraspecific variability matters. Plant and Soil, 411, 359–375.

Ali R S, Poll C, Kandeler E. 2018. Dynamics of soil respiration and microbial communities: Interactive controls of temperature and substrate quality. Soil Biology and Biochemistry, 127, 60–70.

Baumann K, Sanaullah M, Chabbi A, Dignac M F, Bardoux G, Steffens M, Kögel Knabner I, Rumpel C. 2013. Changes in litter chemistry and soil lignin signature during decomposition and stabilisation of ¹³C labelled wheat roots in three subsoil horizons. Soil Biology and Biochemistry, 67, 55–61.

Billings S A, Ballantyne I F. 2013. How interactions between microbial resource demands, soil organic matter stoichiometry, and substrate reactivity determine the direction and magnitude of soil respiratory responses to warming. Global Change Biology, 19, 90–102.

Blagodatskaya E, Kuzyakov Y. 2008. Mechanisms of real and apparent priming effects and their dependence on soil microbial biomass and community structure: Critical review. Biology and Fertility of Soils, 45, 115–131.

Chen G, Fang Y, van Zwieten L, Xuan Y, Tavakkoli E, Wang X, Zhang R. 2021. Priming, stabilization and temperature sensitivity of native SOC is controlled by microbial responses and physicochemical properties of biochar. Soil Biology and Biochemistry, 154, 108139.

Cheng W, Parton W J, Gonzalez-Meler M A, Phillips R, Asao S, McNickle G, Brzostek E, Jastrow J. 2014. Synthesis and modeling perspectives of rhizosphere priming. New Phytologist, 201, 31–44.

Creamer C A, de Menezes A B, Krull E S, Sanderman J, New-ton-Walters R, Farrell M. 2015. Microbial community structure mediates response of soil C decomposition to litter addition and warming. Soil Biology and Biochemistry, 80, 175–188.

Crowther T W,  Todd-Brown K E O, Rowe C W, Wieder W R, Carey J C, Machmuller M B, Snoek B L, Fang S, Zhou G, Allison S D, Blair J M, Bridgham S D, Burton A J, Carrillo Y, Reich P B, Clark J S, Classen A T, Dijkstra F A, Elberling B, Emmett B A, et al. 2016. Quantifying global soil carbon losses in response to warming. Nature, 540, 104–108.

Cui J, Wei L, Zhu Z K, Deng Y W, Chen S, Kuzyakov Y, Ge T D. 2024. Influence of soil N availability on priming effects depending on temperature. Soil Tillage Research, 242, 106163.

Cui J, Zhu Z, Xu X, Liu S, Jones D L, Kuzyakov Y, Shibistova O, Wu J, Ge T. 2020. Carbon and nitrogen recycling from microbial necromass to cope with C:N stoichiometric imbalance by priming. Soil Biology and Biochemistry, 142, 107720.

Cui Y, Fang L, Guo X, Wang X, Zhang Y, Li P, Zhang X. 2018. Ecoenzymatic stoichiometry and microbial nutrient limitation in rhizosphere soil in the arid area of the northern Loess Plateau, China. Soil Biology and Biochemistry, 116, 11–21.

Davis K E R, Sangwan P, Janssen P H. 2011. Acidobacteria, Rubrobacteridae and Chloroflexi are abundant among very slow growing and mini-colony-forming soil bacteria. Environmental Microbiology, 13, 798–805.

Ding J X, He Y H, Yin L M, Huang C J, Xue K, Yan S B, Liu R Q, Wang P, Zhou X H. 2025. Divergent responses of soil positive and negative priming effects to experimental warming. Global Ecology and Biogeography, 34, e70028.

Dong H, Lin J, Lu J, Li L, Yu Z, Kumar A, Zhang Q, Liu D, Chen B. 2022. Priming effects of surface soil organic carbon decreased with warming: A global meta-analysis. Plant and Soil, 500, 233–242.

Dong M Q, Zhao J, Li E, Liu Z J, Guo S B, Zhang Z T, Cui W Q, Yang X G. 2023. Effects of changing climate extremes on maize grain yield in Northeast China. Agronomy, 13, 1050.

Douglas G M, Mafei V J, Zaneveld J, Yurgel S N, Brown J R, Taylor C M, Huttenhower C, Langille M G I. 2019. PICRUSt2: An improved and extensible approach for metagenome inference. BioRxiv, 672295.

Eilers K G, Lauber C L, Knight R, Fierer N. 2010. Shifts in bacterial community structure associated with inputs of low molecular weight carbon compounds to soil. Soil Biology and Biochemistry, 42, 896–903.

Fang Y, Van Zwieten L, Rose M T, Vasileiadis S, Donner E, Vancov T, Rigg J L, Weng Z, Lombi E, Drigo B, Conyers M, Tavakkoli E. 2022. Unraveling microbiomes and functions associated with strategic tillage, stubble, and fertilizer management. Agriculture, Ecosystems & Environment, 323, 107686.

Fanin N, Alavoine G, Bertrand I. 2020. Temporal dynamics of litter quality, soil properties and microbial strategies as main drivers of the priming effect. Geoderma, 377, 114576.

Feng J G, Tang M, Zhu B. 2021. Soil priming effect and its responses to nutrient addition along a tropical forest elevation gradient.  Global Change Biology, 27, 2793-2806.

Fierer N, Bradford M A, Jackson A R B. 2007. Toward an ecological classification of soil bacteria. Ecology, 88, 1354–1364.

Fontaine S, Bardoux G, Abbadie L, Mariotti A. 2004. Carbon input to soil may decrease soil carbon content. Ecology Letters, 7, 314–320.

Fu Y Y, Liu W L, Chen Z Y, Redmile-Gordon M, Liang C, Tang C, Guggenberger G, Yan S B, Yin L M, Peng J J, Zwieten L V, Wang P, Chen J, Kuzyakov Y, Ge T D, Xu J M, Luo Y. 2025. Microbial metabolisms determine soil priming effect induced by organic inputs. Soil Biology and Biochemistry, 209, 109885.

Ghee C, Neilson R, Hallett P D, Robinson D, Paterson E. 2013. Priming of soil organic matter mineralization is intrinsically insensitive to temperature. Soil Biology and Biochemistry, 66, 20–28.

Guo L, Yu Z, Li Y, Xie Z, Wang G, Liu J, Hu X, Wu J, Liu X, Jin J. 2023. Stimulation of primed carbon under climate change corresponds with phosphorus mineralization in the rhizosphere of soybean. Science of the Total Environment, 899, 165580.

Guttières R, Nunan N, Raynaud X, Lacroix G, Barot S, Barré P, Girardin C, Guenet B, Lata J C, Abbadie L. 2021. Temperature and soil management effects on carbon fluxes and priming effect intensity. Soil Biology and Biochemistry, 153, 108103.

Hou Z N, Liu W Z, Xu S Q, Chang S, Zhang X J, Wang R H, Luo D Q, Wang B R, An S S, Qiao Y H, Du Z L, Wei Y Q. 2025. Warming-induced response of microbial diversity and functions regulated microbial necromass and soil multifunctionality. Applied Soil Ecology, 206, 105798.

Huang Q, Wang B R, Shen J K, Xu F J, Li N, Jia P H, Jia Y J, An S S, Amoah I D, Huang Y M. 2024. Shifts in C-degradation genes and microbial metabolic activity with vegetation types affected the surface soil organic carbon pool. Soil Biology and Biochemistry, 192, 109371.

Huo C F, Luo Y Q, Cheng W X. 2017. Rhizosphere priming effect: a meta-analysis. Soil Biology and Biochemistry, 111, 78–84.

Jenkinson D S, Brookes P C, Powlson D S. 2004. Measuring soil microbial biomass. Soil Biology and Biochemistry, 36, 5–7.

Kaiser C, Franklin O, Dieckmann U, Richter A. 2014. Microbial community dynamics alleviate stoichiometric constraints during litter decay. Ecology Letters, 7, 680–690.

Karimi B, Maron P A, Chemidlin-Prevost Boure N, Bernard N, Gilbert D, Ranjard L. 2017. Microbial diversity and ecological networks as indicators of environmental quality. Environmental Chemistry Letters, 15, 265–281.

Khodadad C L M M, Zimmerman A R, Green S J, Uthandi S, Foster J S. 2011. Taxa-specific changes in soil microbial community composition induced by pyrogenic carbon amendments. Soil Biology and Biochemistry, 43, 385–392.

Kuzyakov Y. 2010. Priming effects: Interactions between living and dead organic matter. Soil Biology and Biochemistry, 42, 1363–1371.

Kuzyakov Y, Mason-Jones K. 2018. Viruses in soil: Nano-scale undead drivers of microbial life, biogeochemical turnover and ecosystem functions. Soil Biology and Biochemistry, 127, 305–317.

Lehmann J, Kleber M. 2015. The contentious nature of soil organic matter. Nature, 528, 60–68.

Li H, Yang S, Semenov M V, Yao F, Ye J, Bu R, Ma R, Lin J, Kurganova I, Wang X, Deng Y, Kravchenko I, Jiang Y, Kuzyakov Y. 2021. Temperature sensitivity of SOM decomposition is linked with a K-selected microbial community. Global Change Biology, 27, 2763–2779.

Li L, Zhu-Barker X, Ye R, Doane T A, Horwath W R. 2018. Soil microbial biomass size and soil carbon influence the priming effect from carbon inputs depending on nitrogen availability. Soil Biology and Biochemistry, 119, 41–49.

Li X, Feng J, Zhang Q, Zhu B. 2023. Warming inhibits the priming effect of soil organic carbon mineralization: A meta-analysis. Science of the Total Environment, 904, 166170.

Liang C, Amelung W, Lehmann J, Kastner M. 2019. Quantitative assessment of microbial necromass contribution to soil organic matter. Global Change Biology, 25, 3578–3590.

Liang Y, Cao D, Ma Z, Wu R, Zhang H, Fang Y, Shahbaz M, Liu X J A, Kuzyakov Y, Chen J, Ge T. 2024. Stoichiometry regulates rice straw-induced priming effect: The microbial life strategies. Soil Biology and Biochemistry, 196, 109514.

Lyu L H, Wang C Q, Fan K K, Li J S, Yang T, Gao G F, Sun R, Wang J, Xu X Y, Zhang Y X, Ma Y Y, Zhang J B, Kuzyakov Y, Chu H Y. 2025. Microbial life-history strategies mediate temperature effects on organic carbon pools in black soils. Soil Ecology Letters, 7, 250306.

Ma S, Zhu W, Wang W, Li X, Sheng Z. 2023. Microbial assemblies with distinct trophic strategies drive changes in soil microbial carbon use efficiency along vegetation primary succession in a glacier retreat area of the southeastern Tibetan Plateau. Science of the Total Environment, 867, 161587.

Paul E A. 2007. Soil Microbiology, Ecology, and Biochemistry. Academic Press, New York.

Pausch J, Holz M, Zhu B, Cheng W X. 2024. Rhizosphere priming promotes plant nitrogen acquisition by microbial necromass recycling. Plant, Cell & Environment, 47, 1987-1996.

Pingthaisong W, Blagodatsky S, Vityakon P, Cadisch G. 2024. Mixing plant residues of different quality reduces priming effect and contributes to soil carbon retention. Soil Biology and Biochemistry, 188, 109242.

Razanamalala K, Razafimbelo T, Maron P A, Ranjard L, Chemidlin N, Lelievre M, Dequiedt S, Ramaroson V H, Marsden C, Becquer T, Trap J, Blanchart E, Bernard L. 2018. Soil microbial diversity drives the priming effect along climate gradients: A case study in Madagascar. ISME Journal, 12, 451–462.

Reji L, Darnajoux R, Zhang X N. 2024. A genomic view of environmental and life history controls on microbial nitrogen acquisition strategies. Environmental Microbiology Reports, 16, e13220.

Ren C, Wang J, Bastida F, Delgado-Baquerizo M, Yang Y, Wang J, Zhong Z, Zhou Z, Zhang S, Guo Y, Zhou S, Wei G, Han X, Yang G, Zhao F. 2022. Microbial traits determine soil C emission in response to fresh carbon inputs in forests across biomes. Global Change Biology, 28, 1516–1528.

Ren T, Huang X, Sun H, Xie H, Jia Z, Wang J. 2025. Comparison on responses of black soil microorganisms to exogenous carbon and nitrogen addition under no-tillage with straw mulching and combined application of organic and inorganic fertilizers. Acta Microbiologica Sinica, 65, 3348-3364. (in Chinese)

Song X, Liu X, Liang G, Li S, Li J, Zhang M, Zheng F, Ding W, Wu X, Wu H. 2022. Positive priming effect explained by microbial nitrogen mining and stoichiometric decomposition at different stages. Soil Biology and Biochemistry, 175, 108852.

Tan F, Cheng J, Zhang Y, Jiang X, Liu Y. 2022. Genomics analysis and degradation characteristics of lignin by Streptomyces thermocarboxydus strain DF3-3. Biotechnology for Biofuels and Bioproducts, 15, 1–15.

Tao X Y, Yang Z F, Feng J J, Jian S Y,Yang Y F, Bates C T, Wang G S, Guo X, Ning D L, Kempher M L, Liu X J A, Ouyang Y, Han S, Wu L W, Zeng Y F, Kuang J L, Zhang Y, Zhou X S, Shi Z, Qin W, et al. 2024. Experimental warming accelerates positive soil priming in a temperate grassland ecosystem. Nature Communications, 15, 1178.

Tian Q, Jiang Q, Zhao R, Wu Y, Lin Q, Zhao X, Tang Z, Liu F. 2023. Microbial properties control soil priming and exogenous carbon incorporation along an elevation gradient. Geoderma, 431, 116343.

Tong Y X, Liu J G, Li X L, Sun J, Herzberger A, Wei D, Zhang W F, Dou Z X, Zhang F S. 2017. Cropping system conversion led to organic carbon change in China’s Mollisols regions. Scientific Report, 7, 18064.

Vance E D, Brookes P C, Jenkinson D S. 1987. An extraction method for measuring soil microbial biomass C. Soil Biology and Biochemistry, 19, 703–707.

Wagg C, Schlaeppi K, Banerjee S, Kuramae E E, van der Heijden M G A. 2019. Fungalbacterial diversity and microbiome complexity predict ecosystem functioning. Nature Communications, 10, 4841.

Wang J, Pan F, Soininen J, Heino J, Shen J. 2016. Nutrient enrichment modifies temperature biodiversity relationships in large-scale field experiments. Nature Communications, 7, 13960.

Wang J, Yao H Y, Zhang X. 2024. The effect of the ¹³C abundance of soil microbial DNA on identifying labelled fractions after ultracentrifugation. Applied Microbiology and Biotechnology, 108, 318.

Xu Q, Jin J, Wang X, Armstrong R, Tang C. 2019. Susceptibility of soil organic carbon to priming after long-term CO₂ fumigation is mediated by soil texture. Science of the Total Environment, 657, 1112–1120.

Yanni S F, Diochon A, Helgason B L, Ellert B H, Gregorich E G. 2018. Temperature response of plant residue and soil organic matter decomposition in soil from different depths. European Journal of Soil Science, 69, 325–335.

You M, He P, Dai S, Burger M, Li L J. 2021. Priming effect of stable C pool in soil and its temperature sensitivity. Geoderma, 401, 115216.

Yu X Y, Ma Y Y. 2022. Spatial and temporal analysis of extreme climate events over Northeast China. Atmosphere, 13, 1197.

Yu Z Y, Ling L, Singh BP, Luo Y, Xu J M. 2020. Gain in carbon: Deciphering the abiotic and biotic mechanisms of biochar-induced negative priming effects in contrasting soils. Science of the Total Environment, 746, 141057.

Zhang H, Chang D, Zhu Z, Meng C, Wang K. 2024. Soil priming effects and involved microbial community along salt gradients. Biogeosciences, 21, 1–11.

Zhao F, Wang J, Li Y, Xu X, He L, Wang J, Ren C, Guo Y. 2022. Microbial functional genes driving the positive priming effect in forest soils along an elevation gradient. Soil Biology and Biochemistry, 165, 108498.

Zhao J, Xie X, Jiang Y, Li J, Fu Q, Qiu Y, Fu X, Yao Z, Dai Z, Qiu Y. 2024. Effects of simulated warming on soil microbial community diversity and composition across diverse ecosystems. Science of the Total Environment, 911, 168793.

Zheng Y, Wang X, Hayden H, Franks A, Shindler A, Liu Y, Clark G, Jin J, Tang C. 2023. Soil types differ in the temporal response of the priming effect to N addition: A study on microbial mechanisms. Biology and Fertility of Soils, 59, 233–247.

Zhong Z K, Li W J, Lu X Q, Gu Y Q, Wu S J, Shen Z Y, Han X H, Yang G H, Ren C J. 2020. Adaptive pathways of soil microorganisms to stoichiometric imbalances regulate microbial respiration following afforestation in the Loess Plateau, China. Soil Biology and Biochemistry, 151, 108048.

Zhou J, Liu Y, Liu C Y, Zamanian K, Feng W H, Steiner S K, Shi L L, Guillaume T, Kumar A. 2024. Necromass responses to warming: A faster microbial turnover in favor of soil carbon stabilisation. Science of the Total Environment, 954,176651.

Zhou S, Wang J, Chen L, Wang J, Zhao F. 2022. Microbial community structure and functional genes drive soil priming effect following afforestation. Science of the Total Environment, 825, 153925.

Zhu Z, Fang Y, Liang Y, Li Y, Liu S, Li Y, Li B, Gao W, Yuan H, Kuzyakov Y, Wu J, Richter A, Ge T. 2022. Stoichiometric regulation of priming effects and soil carbon balance by microbial life strategies. Soil Biology and Biochemistry, 169, 108669.