Ali U, Lu S, Fadlalla T, Iqbal S, Yue H, Yang B, Hong Y, Wang X, Guo L. 2022. The functions of phospholipases and their hydrolysis products in plant growth, development and stress responses. Progress in Lipid Research, 86, 101158.

Bailey A P, Koster G, Guillermier C, Hirst E M, MacRae J I, Lechene C P, Postle A D, Gould A P. 2015. Antioxidant role for lipid droplets in a stem cell niche of drosophila. Cell, 163, 340–353.

Bates P D. 2016. Understanding the control of acyl flux through the lipid metabolic network of plant oil biosynthesis. Biochimica et Biophysica Acta, 1861, 1214–1225.

Bell E, Creelman R A, Mullet J E. 1995. A chloroplast lipoxygenase is required for wound-induced jasmonic acid accumulation in Arabidopsis. Proceedings of the National Academy of Sciences of the United States of America, 92, 8675–8679.

Benning C. 2009. Mechanisms of lipid transport involved in organelle biogenesis in plant cells. Annual Review of Cell and Developmental Biology, 25, 71–91.

Block M A, Dorne A J, Joyard J, Douce R. 1983. Preparation and characterization of membrane fractions enriched in outer and inner envelope membranes from spinach chloroplasts. I. Electrophoretic and immunochemical analyses. Journal of Biological Chemistry, 258, 13273–13280.

Browse J, Warwick N, Somerville C R, Slack C R. 1986. Fluxes through the prokaryotic and eukaryotic pathways of lipid synthesis in the '16:3' plant Arabidopsis thaliana. The Biochemical Journal, 235, 25–31.

Caldelari D, Wang G, Farmer E E, Dong X. 2011. Arabidopsis lox3 lox4 double mutants are male sterile and defective in global proliferative arrest. Plant Molecular Biology, 75, 25–33.

Cheng X Q, Zhang X Y, Xue F, Zhu S H, Li Y J, Zhu Q H, Liu F, Sun J. 2020. Characterization and transcriptome analysis of a dominant genic male sterile cotton mutant. BMC Plant Biology, 20, 312.

Ding Y, Ma Y, Liu N, Xu J, Hu Q, Li Y, Wu Y, Xie S, Zhu L, Min L, Zhang X. 2017. microRNAs involved in auxin signalling modulate male sterility under high-temperature stress in cotton (Gossypium hirsutum). The Plant Journal, 91, 977–994.

Fan J, Yan C, Xu C. 2013. Phospholipid: Diacylglycerol acyltransferase-mediated triacylglycerol biosynthesis is crucial for protection against fatty acid-induced cell death in growing tissues of Arabidopsis. The Plant Journal, 76, 930–942.

Fu W, Shen Y, Hao J, Wu J, Ke L, Wu C, Huang K, Luo B, Xu M, Cheng X, Zhou X, Sun J, Xing C, Sun Y. 2015. Acyl-CoA N-acyltransferase influences fertility by regulating lipid metabolism and jasmonic acid biogenesis in cotton. Scientific Reports, 5, 11790.

Gao Y, Chen G, Weselake R J. 2014. A rapid nile red fluorescence-based method for triacylglycerol content in microspore-derived cell suspension cultures of Brassica napus. Lipids, 49, 1161–1168.

Han G S, Carman G M. 2010. Characterization of the human LPIN1-encoded phosphatidate phosphatase isoforms. The Journal of Biological Chemistry, 285, 14628–14638.

He M, Ding N Z. 2020. Plant unsaturated fatty acids: Multiple roles in stress response. Frontiers in Plant Science, 11, 562785.

Heinz E, Roughan P G. 1983. Similarities and differences in lipid metabolism of chloroplasts isolated from 18:3 and 16:3 plants. Plant Physiology, 72, 273–279.

Hyun Y, Choi S, Hwang H J, Yu J, Nam S J, Ko J, Park J Y, Seo Y S, Kim E Y, Ryu S B, Kim W T, Lee Y H, Kang H, Lee I. 2008. Cooperation and functional diversification of two closely related galactolipase genes for jasmonate biosynthesis. Developmental Cell, 14, 183–192.

Ishiguro S, Kawai-Oda A, Ueda J, Nishida I, Okada K. 2001. The DEFECTIVE IN ANTHER DEHISCIENCE1 gene encodes a novel phospholipase A1 catalyzing the initial step of jasmonic acid biosynthesis, which synchronizes pollen maturation, anther dehiscence, and flower opening in Arabidopsis. The Plant Cell, 13, 2191–2209.

Jiang P, Zhang X, Zhu Y, Zhu W, Xie H, Wang X. 2007. Metabolism of reactive oxygen species in cotton cytoplasmic male sterility and its restoration. Plant Cell Reports, 26, 1627–1634.

Khan A H, Min L, Ma Y, Wu Y, Ding Y, Li Y, Xie S, Ullah A, Shaban M, Manghwar H, Shahid M, Zhao Y, Wang C, Zhang X. 2020. High day and night temperatures distinctively disrupt fatty acid and jasmonic acid metabolism, inducing male sterility in cotton. Journal of Experimental Botany, 71, 6128–6141.

Kohlwein S D. 2010. Triacylglycerol homeostasis: Insights from yeast. The Journal of Biological Chemistry, 285, 15663–15667.

Lavell A A, Benning C. 2019. Cellular organization and regulation of plant glycerolipid metabolism. Plant Cell Physiology, 60, 1176–1183.

Li L O, Klett E L, Coleman R A. 2010. Acyl-CoA synthesis, lipid metabolism and lipotoxicity. Biochimica et Biophysica Acta, 1801, 246–251.

Li M, Hong Y, Wang X. 2009. Phospholipase D- and phosphatidic acid-mediated signaling in plants. Biochimica et Biophysica Acta, 1791, 927–935.

Li-Beisson Y, Shorrosh B, Beisson F, Andersson M X, Arondel V, Bates P D, Baud S, Bird D, Debono A, Durrett T P, Franke R B, Graham I A, Katayama K, Kelly A A, Larson T, Markham J E, Miquel M, Molina I, Nishida I, Rowland O, et al. 2013. Acyl-lipid metabolism. The Arabidopsis Book, 11, e0161.

Liu F, Ma L, Wang Y, Li Y, Zhang X, Xue F, Nie X, Zhu Q, Sun J. 2019. GhFAD2-3 is required for anther development in Gossypium hirsutum. BMC Plant Biology, 19, 393.

Liu F, Zhao Y P, Zhu H G, Zhu Q H, Sun J. 2017. Simultaneous silencing of GhFAD2-1 and GhFATB enhances the quality of cottonseed oil with high oleic acid. Journal of Plant Physiology, 215, 132–139.

Lock Y Y, Snyder C L, Zhu W, Siloto R M, Weselake R J, Shah S. 2009. Antisense suppression of type 1 diacylglycerol acyltransferase adversely affects plant development in Brassica napus. Physiologia Plantarum, 137, 61–71.

Lohden I, Frentzen M. 1988. Role of plastidial acyl-acyl carrier protein: Glycerol 3-phosphate acyltransferase and acyl-acyl carrier protein hydrolase in channelling the acyl flux through the prokaryotic and eukaryotic pathway. Planta, 176, 506–512.

Ma H, Wu Y, Lv R, Chi H, Zhao Y, Li Y, Liu H, Ma Y, Zhu L, Guo X, Kong J, Wu J, Xing C, Zhang X, Min L. 2022. Cytochrome P450 mono-oxygenase CYP703A2 plays a central role in sporopollenin formation and ms5ms6 fertility in cotton. Journal of Integrative Plant Biology, 64, 2009–2025.

von Malek B, van der Graaff E, Schneitz K, Keller B. 2002. The Arabidopsis male-sterile mutant dde2-2 is defective in the ALLENE OXIDE SYNTHASE gene encoding one of the key enzymes of the jasmonic acid biosynthesis pathway. Planta, 216, 187–192.

Maraschin F, Kulcheski F R, Segatto A, Trenz T S, Barrientos-Diaz O, Margis-Pinheiro M, Margis R, Turchetto-Zolet A C. 2019. Enzymes of glycerol-3-phosphate pathway in triacylglycerol synthesis in plants: Function, biotechnological application and evolution. Progress in Lipid Research, 73, 46–64.

McConn M, Browse J. 1996. The critical requirement for linolenic acid is pollen development, not photosynthesis, in an Arabidopsis mutant. The Plant Cell, 8, 403–416.

Min L, Li Y, Hu Q, Zhu L, Gao W, Wu Y, Ding Y, Liu S, Yang X, Zhang X. 2014. Sugar and auxin signaling pathways respond to high-temperature stress during anther development as revealed by transcript profiling analysis in cotton. Plant Physiology, 164, 1293–1308.

Mittler R. 2002. Oxidative stress, antioxidants and stress tolerance. Trends in Plant Science, 7, 405–410.

Ohlrogge J, Browse J. 1995. Lipid biosynthesis. The Plant Cell, 7, 957–970.

Park J H, Halitschke R, Kim H B, Baldwin I T, Feldmann K A, Feyereisen R. 2002. A knock-out mutation in allene oxide synthase results in male sterility and defective wound signal transduction in Arabidopsis due to a block in jasmonic acid biosynthesis. The Plant Journal, 31, 1–12.

Piffanelli P, Ross J H, Murphy D. 1998. Biogenesis and function of the lipidic structures of pollen grains. Sexual Plant Reproduction, 11, 65–80.

Pressman E, Shaked R, Shen S, Altahan L, Firon N. 2012. Variations in carbohydrate content and sucrose-metabolizing enzymes in tomato (Solanum lycopersicum L.) stamen parts during pollen maturation. American Journal of Plant Sciences, 3, 252–260.

Rong H, Yang W, Xie T, Wang Y, Wang X, Jiang J, Wang Y. 2022. Transcriptional profiling between yellow- and black-seeded Brassica napus reveals molecular modulations on flavonoid and fatty acid content. Journal of Integrative Agriculture, 21, 2211–2226.

Sawhney V K, Shukla S A. 1994. Male sterility in flowering plants: Are plant growth substances involved. American Journal of Botany, 81, 1640–1647.

Schilmiller A L, Koo A J, Howe G A. 2007. Functional diversification of acyl-coenzyme A oxidases in jasmonic acid biosynthesis and action. Plant Physiology, 143, 812–824.

Stenzel I, Hause B, Miersch O, Kurz T, Maucher H, Weichert H, Ziegler J, Feussner I, Wasternack C. 2003. Jasmonate biosynthesis and the allene oxide cyclase family of Arabidopsis thaliana. Plant Molecular Biology, 51, 895–911.

Stintzi A, Browse J. 2000. The Arabidopsis male-sterile mutant, opr3, lacks the 12-oxophytodienoic acid reductase required for jasmonate synthesis. Proceedings of the National Academy of Sciences of the United States of America, 97, 10625–10630.

Taguchi R, Ishikawa M. 2010. Precise and global identification of phospholipid molecular species by an Orbitrap mass spectrometer and automated search engine Lipid Search. Journal of Chromatography A, 1217, 4229–4239.

Vance J E, Vance D E. 2004. Phospholipid biosynthesis in mammalian cells. Biochemistry and Cell Biology, 82, 113–128.

Wan X, Wu S, Li Z, An X, Tian Y. 2020. Lipid metabolism: Critical roles in male fertility and other aspects of reproductive development in plants. Molecular Plant, 13, 955–983.

Wasternack C, Hause B. 2013. Jasmonates: Biosynthesis, perception, signal transduction and action in plant stress response, growth and development. Annals of Botany, 111, 1021–1058.

Wasternack C, Song S. 2017. Jasmonates: Biosynthesis, metabolism, and signaling by proteins activating and repressing transcription. Journal of Experimental Botany, 68, 1303–1321.

Weber H. 2002. Fatty acid-derived signals in plants. Trends in Plant Science, 7, 217–224.

Wu Y, Li Y, Li Y, Ma Y, Zhao Y, Wang C, Chi H, Chen M, Ding Y, Guo X, Min L, Zhang X. 2019. Proteomic analysis reveals that sugar and fatty acid metabolisms play a central role in sterility of the male-sterile line 1355A of cotton. The Journal of Biological Chemistry, 294, 7057–7067.

Wu Y, Min L, Wu Z, Yang L, Zhu L, Yang X, Yuan D, Guo X, Zhang X. 2015. Defective pollen wall contributes to male sterility in the male sterile line 1355A of cotton. Scientific Reports, 5, 9608.

Xie H T, Wan Z Y, Li S, Zhang Y. 2014. Spatiotemporal production of reactive oxygen species by NADPH oxidase is critical for tapetal programmed cell death and pollen development in Arabidopsis. The Plant Cell, 26, 2007–2023.

Xu C, Fan J, Shanklin J. 2020. Metabolic and functional connections between cytoplasmic and chloroplast triacylglycerol storage. Progress in Lipid Research, 80, 101069.

Yang J. 2001. Hormonal changes in the grains of rice subjected to water stress during grain fifilling. Plant Physiology, 127, 315–323.

Yen C L, Stone S J, Koliwad S, Harris C, Farese Jr R V. 2008. Thematic review series: Glycerolipids. DGAT enzymes and triacylglycerol biosynthesis. Journal of Lipid Research, 49, 2283–2301.

Yu C W, Lin Y T, Li H M. 2020. Increased ratio of galactolipid MGDG: DGDG induces jasmonic acid overproduction and changes chloroplast shape. The New Phytologist, 228, 1327–1335.

Yurchenko O P, Park S, Ilut D C, Inmon J J, Millhollon J C, Liechty Z, Page J T, Jenks M A, Chapman K D, Udall J A, Gore M A, Dyer J M. 2014. Genome-wide analysis of the omega-3 fatty acid desaturase gene family in Gossypium. BMC Plant Biology, 14, 312.

Zegarlińska J, Piaścik M, Sikorski A F, Czogalla A. 2018. Phosphatidic acid-a simple phospholipid with multiple faces. Acta Biochimica Polonica, 65, 163–171.

Zhang T, Hu Y, Jiang W, Fang L, Guan X, Chen J, Zhang J, Saski C A, Scheffler B E, Stelly D M, Hulse-Kemp A M, Wan Q, Liu B, Liu C, Wang S, Pan M, Wang Y, Wang D, Ye W, Chang L, et al. 2015. Sequencing of allotetraploid cotton (Gossypium hirsutum L. acc. TM-1) provides a resource for fiber improvement. Nature Biotechnology, 33, 531–537.

Zhang Y, Zhu H, Zhang Q, Li M, Yan M, Wang R, Wang L, Welti R, Zhang W, Wang X. 2009. Phospholipase dalpha1 and phosphatidic acid regulate NADPH oxidase activity and production of reactive oxygen species in ABA-mediated stomatal closure in Arabidopsis. The Plant Cell, 21, 2357–2377.

Zhu Y, Wang K, Wu C, Zhao Y, Yin X, Zhang B, Grierson D, Chen K, Xu C. 2019. Effect of ethylene on cell wall and lipid metabolism during alleviation of postharvest chilling injury in peach. Cells, 8, 1612.

Zhu Y, Yuan G, Jia S, An G, Li W, Sun D, Liu J. 2022. Transcriptomic profiling of watermelon (Citrullus lanatus) provides insights into male flowers development. Journal of Integrative Agriculture, 21, 407–421.