Akhberdi O, Zhang Q, Wang D, Wang H, Hao X, Liu Y, Wei D S, Zhu X D. 2018. Distinct Roles of Velvet complex in the development, stress tolerance, and secondary metabolism in Pestalotiopsis microspora, a taxol producer. Gene, 9, 164.

Bai G, Shaner G E. 1996. Variation in Fusarium graminearum and cultivar resistance to wheat scab. Plant Disease, 80, 975-979.

Bayram O, Braus G H, Fischer R, Rodriguez-Romero J. 2010. Spotlight on Aspergillus nidulans photosensory systems. Fungal Genetics and Biology, 47, 900-908.

Bayram O, Krappmann S, Ni M, Bok J W, Helmstaedt K, Valerius O, Braus-Stromeyer S, Kwon N J, Keller N P, Yu J H, Braus G H. 2008. VelB/VeA/LaeA complex coordinates light signal with fungal development and secondary metabolism. Science, 320, 1504-1506.

Bazafkan H, Dattenbock C, Stappler E, Beier S, Schmoll M. 2017. Interrelationships of VEL1 and ENV1 in light response and development in Trichoderma reesei. PLoS One, 12, e0175946.

Bowers J H, Locke J C. 2000. Effect of botanical extracts on the population density of Fusarium oxysporum in soil and control of Fusarium Wilt in the greenhouse. Plant Disease, 84, 300-305.

Brown D, Lee S H, Kim L, Ryu J G, Lee S, Seo Y, Kim Y H, Busman M, Yun S H, Proctor R H, Lee T. 2014. Identification of a 12-gene Fusaric Acid biosynthetic gene cluster in Fusarium species through comparative and functional genomics. Molecular Plant-Microbe Interactions, 28, 319-332.

Brown A, Budge S, Kaloriti D, Tillmann A, Jacobsen M, Yin Z K, Ene I V, Bohovych I, Sandai D, Kastora S, Potrykus J, Ballou E R, Childers D S, Shahana S, Leach M D. 2014. Stress adaptation in a pathogenic fungus. Journal of Experimental Biology. 217, 144-155.

Butchko R E, Brown D , Busman M, Tudzynski B, Wiemann P. 2012. LAE1 regulates expression of multiple secondary metabolite gene clusters in Fusarium verticillioides. Fungal Genetics and Biology, 49, 602-612.

Choi Y E, Goodwin S. 2011. MVE1, Encoding the velvet gene product homolog in Mycosphaerella graminicola, is associated with aerial mycelium formation, melanin biosynthesis, hyphal swelling, and light signaling. Applied and Environmental Microbiology, 77, 942-953.

Ding S, Zhou D, Wei H, Wu S, Xie B. 2021. Alleviating soil degradation caused by watermelon continuous cropping obstacle: Application of urban waste compost. Chemosphere, 262, 128387.

Dong X, Ling N, Wang M, Shen Q, Guo S. 2012. Fusaric acid is a crucial factor in the disturbance of leaf water imbalance in Fusarium-infected banana plants. Plant Physiology and Biochemistry, 60, 171-179.

Duan Y, Ge C, Liu S, Wang J, Zhou M. 2013. A two-component histidine kinase Shk1 controls stress response, sclerotial formation and fungicide resistance in Sclerotinia sclerotiorum. Molecular Plant Pathology, 14, 708-718.

Estiarte N, Lawrence C B, Sanchis V, Ramos A J, Crespo-Sempere A. 2016. LAEA and VEA are involved in growth morphology, asexual development, and mycotoxin production in Alternaria alternata. International Journal of Food Microbiology, 238, 153-164.

Fravel D, Olivain C, Alabouvette C. 2003. Fusarium oxysporum and its biocontrol. New Phytologist, 157, 493-502.

Fisher M C, Henk D A, Briggs C J, Brownstein J S, Madoff L C, McCraw S L, Gurr S J. 2012. Emerging fungal threats to animal, plant and ecosystem health. Nature, 484, 186-194.

Gies, H, Sondergaard T E, Sørensen J L. 2013. The AreA transcription factor in Fusarium graminearum regulates the use of some nonpreferred nitrogen sources and secondary metabolite production. Fungal Biology, 117, 814-821.

Guo S, Zhang J, Sun H, Salse J, Lucas WJ, Zhang H, Zheng Y, Mao L, Ren Y, Wang Z, Min J, Guo X, Murat F, Ham B K, Zhang Z, Gao S, Huang M, Xu Y, Zhong S, Bombarely A, Mueller L A, Zhao H, He H, Zhang Y, Zhang Z, Huang S, Tan T, Pang E, Lin K, Hu Q, Kuang H, Ni P, Wang B, Liu J, Kou Q, Hou W, Zou X, Jiang J, Gong G, Klee K, Schoof H, Huang Y, Hu X, Dong S, Liang D, Wang J, Wu K, Xia Y, Zhao X, Zheng Z, Xing M, Liang X, Huang B, Lv T, Wang J, Yin Y, Yi H, Li R, Wu M, Levi A, Zhang X, Giovannoni J J, Wang J, Li Y, Fei Z, Xu Y. 2013. The draft genome of watermelon Citrullus lanatus. and resequencing of 20 diverse accessions. Nature Genetics, 45, 51-58.

He W J, Zhang L, Yi S Y, Tang X L, Yuan Q S, Guo M W, Wu A B, Qu B, Li H P, Liao Y C. 2017. An aldo-keto reductase is responsible for Fusarium toxin-degrading activity in a soil Sphingomonas strain. Scientific Reports, 7, 9549.

Hofer A M, Harting R, Aßmann N F, Gerke J, Schmitt K, Starke J, Bayram O, Tran V T, Valerius O, Braus-Stromeyer S A, Braus G H. 2021. The velvet protein VEL1 controls initial plant root colonization and conidia formation for xylem distribution in Verticillium wilt. PLoS Genetics, 17, e1009434.

Hua g k, Wang L, Chen J, Ji P S. 2019. Biological control of Fusarium wilt on watermelon by fluorescent pseudomonads. Biocontrol Science and Technology, 30, 1-16.

Jiang C, Zhang C, Wu C, Sun P, Hou R, Liu H, Wang C F, Xu J R. 2016. TRI6 and TRI10 play different roles in the regulation of deoxynivalenol DON. production by cAMP signalling in Fusarium graminearum. Environmental Microbiology, 18, 3689-3701.

Karimi-Aghcheh R, Bok J W, Phatale P A, Smith K M, Baker S E, Lichius A, Omann M, Zeilinger S, Seiboth B, Rhee C, Keller N P, Freitag M, Kubicek C P. 2013. Functional Analyses of Trichoderma reesei LAE1 Reveal Conserved and Contrasting Roles of This Regulator. Genes, 3, 369-378.

Keinath A P, Wechter W P, Rutter W B, Agudelo P A. 2019. Cucurbit Rootstocks Resistant to Fusarium oxysporum f. sp. niveum Remain Resistant When Coinfected by Meloidogyne incognita in the Field. Plant Disease, 103, 1383-1390.

Krappmann S, Bayram O, Braus G H. 2005. Deletion and allelic exchange of the Aspergillus fumigatus VEA locus via a novel recyclable marker module. Eukaryotic Cell, 4, 1298-1307.

Kuang Y, Scherlach K, Hertweck C, Yang S, Sampietro D A, Karlovsky P. 2022. Fusaric acid detoxification: a strategy of Gliocladium roseum involved in its antagonism against Fusarium verticillioides. Mycotoxin Research, 38, 13-25.

Levin D E. 2005. Cell wall integrity signaling in Saccharomyces cerevisiae. Microbiology and Molecular Biology Reviews. 69, 262–291.

Lin, C H and Chung K R. 2010. Specialized and shared functions of the histidine kinase and HOG1 MAP kinase-mediated signaling pathways in Alternaria alternata, a filamentous fungal pathogen of citrus. Fungal Genetics and Biology. 47, 818-827.

Li H, Yuan G, Zhu C, Zhao T, Ruimin Z, Wang X L, Yang J Q, Ma Jian X, Zhang Y, Zhang X. 2019. Soil fumigation with ammonium bicarbonate or metam sodium under high temperature alleviates continuous cropping-induced Fusarium wilt in watermelon. Scientia Horticulturae, 246, 979-986.

Liu K, Dong Y, Wang F, Jiang B, Wang M, Fang X. 2016. Regulation of cellulase expression, sporulation, and morphogenesis by velvet family proteins in Trichoderma reesei. Applied Microbiology and Biotechnology, 100, 769-779.

Liu S, Li J, Zhang Y, Liu N, Viljoen A, Mostert D, Mostert D, Zuo C, Hu C H, Bi F C, Gao H J, Sheng O, Deng G M, Yang Q S, Dong T, Dou T X, Yi G J, Ma L J, Li C Y. 2020. Fusaric acid instigates the invasion of banana by Fusarium oxysporum f. sp. cubense TR4. New Phytologist, 225, 913-929.

Livak K.J, Schmittgen T D. 2001. Analysis of relative gene expression data using real-time quantitative PCR and the 2 -Delta Delta C T. Method. Methods, 25, 402-408.

Lopez-Berges M, Hera C, Sulyok M, Schäfer K, Capilla J, Guarro J, Pietro A D. 2012. The velvet complex governs mycotoxin production and virulence of Fusarium oxysporum on plant and mammalian hosts. Molecular microbiology, 87, 49-65.

Ma L J, van der Does H C, Borkovich K A, Coleman J J, Daboussi M J, Di Pietro A, Dufresne M, Freitag M, Grabherr M, Henrissat B, Houterman P M, Kang S, Shim W B, Woloshuk C, Xie X, Xu J R, Antoniw J, Baker S E, Bluhm B H, Breakspear A, Brown D W, Butchko R A, Chapman S, Coulson R, Coutinho P M, Danchin E G, Diener A, Gale L R, Gardiner D M, Goff S, Hammond-Kosack K E, Hilburn K, Hua-Van A, Jonkers W, Kazan K, Kodira C D, Koehrsen M, Kumar L, Lee Y H, Li L, Manners J M, Miranda-Saavedra D, Mukherjee M, Park G, Park J, Park S Y, Proctor R H, Regev A, Ruiz-Roldan M C, Sain D, Sakthikumar S, Sykes S, Schwartz D C, Turgeon B G, Wapinski I, Yoder O, Young S, Zeng Q, Zhou S, Galagan J, Cuomo C A, Kistler H C, Rep M. 2010. Comparative genomics reveals mobile pathogenicity chromosomes in Fusarium. Nature, 464, 367-373.

Mamur S, Ünal F, Yılmaz S, Erikel E, Yuzbasioglu D. 2020. Evaluation of the cytotoxic and genotoxic effects of mycotoxin fusaric acid. Drug and Chemical Toxicology, 43, 149-157.

Mateo E M, Tarazona A, Aznar R, Mateo F. 2023. Exploring the impact of lactic acid bacteria on the biocontrol of toxigenic Fusarium spp. and their main mycotoxins. International Journal of Food Microbiology, 387, 110054.

Mukherjee P, Kenerley C. 2010. Regulation of Morphogenesis and Biocontrol Properties in Trichoderma virens by a Velvet Protein, Vel1. Applied and Environmental Microbiology, 76, 2345-2352.

Niehaus E M, Von Bargen K, Espino J, Pfannmuller A, Humpf H U, Tudzynski B. 2014. Characterization of the fusaric acid gene cluster in Fusarium fujikuroi. Applied microbiology and biotechnology, 98, 1749-1762.

Pal S, Eguru S, Reddy D C, L, Dayandhi E. 2023. QTL-seq identifies and QTL mapping refines genomic regions conferring resistance to Fusarium oxysporum f. sp. niveum race 2 in cultivated watermelon [Citrullus lanatus (Thunb.) Matsum & Nakai]. Scientia Horticulturae, 319, 112180.

Perchepied L, Pitrat M. 2005. Polygenic Inheritance of Partial Resistance to Fusarium oxysporum f. sp. melonis Race 1.2 in Melon. Phytopathology, 94, 1331-1336.

Phasha M M, Wingfield B D, Wingfield M J, Coetzee M P A, Hammerbacher A, Steenkamp E T. 2021. Deciphering the effect of FUB1 disruption on fusaric acid production and pathogenicity in Fusarium circinatum. Fungal Biology, 125, 1036-1047.

Inoue I, Namiki F, Tsuge T. 2002. Plant Colonization by the vascular wilt fungus Fusarium oxysporum requires FOW1, a gene encoding a mitochondrial protein. The Plant Cell, 14, 1869-1883.

Karimi-Aghcheh R, Bok J W, Phatale P A, Smith K M, Baker S E, Lichius A. 2013. Functional Analyses of Trichoderma reesei LAE1 Reveal Conserved and Contrasting Roles of This Regulator. G3-Genes Genomes Genetics, 3, 369-378.

Sarikaya-Bayram O, Bayram O, Valerius O, Park H S, Irniger S, Gerke J, Ni M, Han K H, Yu J H, Braus G H. 2010. LAEA control of velvet family regulatory proteins for light-dependent development and fungal cell-type specificity. PLoS Genetics, 6, e1001226.

Schumacher J, Pradier J M, Simon A, Traeger S, Moraga J, Collado I G, Viaud M, Tudzynski B. 2012. Natural variation in the Velvet gene bcvel1 affects virulence and light-dependent differentiation in Botrytis cinerea. PLoS One, 7, e47840.

Schumacher J, Simon A, Cohrs K, Traeger S, Porquier A, Dalmais B, Viaud M, Tudzynski B. 2015. The Velvet Complex in the Gray Mold Fungus Botrytis cinerea : Impact of BcLAE1 on Differentiation, Secondary Metabolism, and Virulence. Molecular Plant-Microbe Interactions, 28, 659-674.

Segorbe D, Pietro A D, Perez N E, Turra D. 2017. Three Fusarium oxysporum mitogen-activated protein kinases (MAPKs) have distinct and complementary roles in stress adaptation and cross-kingdom pathogenicity. Molecular Plant Pathology. 18, 912-924.

Singh V K, Singh H B, Upadhyay R S. 2017. Role of fusaric acid in the development of 'Fusarium wilt' symptoms in tomato: Physiological, biochemical and proteomic perspectives. Plant Physiology and Biochemistry, 118, 320-332.

Tamura K, Dudley J, Nei M, Kumar S. 2007. MEGA4: Molecular Evolutionary Genetics Analysis  MEGA. Software Version 4.0. Molecular Biology and Evolution, 24, 1596-1599.

Tang G, Chen A, Dawood D H, Liang J, Chen Y, Ma Z. 2020. Capping proteins regulate fungal development, DON-toxisome formation and virulence in Fusarium graminearum. Molecular Plant Pathology, 21, 173-187.

Tran M T, Ameye M, Thi-Kim Phan L, Devlieghere F, De Saeger S, Eeckhout M, Audenaert K. 2021. Impact of ethnic pre-harvest practices on the occurrence of Fusarium verticillioides and fumonisin B1 in maize fields from Vietnam. Food Control, 120, 107567.

Tudzynski P, Sharon A. 2003. Fungal Pathogenicity Genes. Applied Mycology and Biotechnology, 3, 187-212.

Van T N , Schafer W, Bormann J. 2012. The stress-activated protein kinase FgOS-2 is a key regulator in the life cycle of the cereal pathogen Fusarium graminearum. Molecular Plant, 25, 1142-1156.

Velluti A, Marín S, Bettucci L, Ramos A J, Sanchis V. 2000. The effect of fungal competition on colonization of maize grain by Fusarium moniliforme, F. proliferatum and F. graminearum and on fumonisin B1 and zearalenone formation. International Journal of Food Microbiology, 59, 59-66.

Wang W, Wu D, Pan H, Turgeon B G. 2014. VEL2 and VOS1 hold essential roles in ascospore and asexual spore development of the heterothallic maize pathogen Cochliobolus heterostrophus. Fungal Genetics and Biology, 70, 113-124.

Wang Y, Duan Y, Wang J, Zhou M. 2015. A new point mutation in the iron-sulfur subunit of succinate dehydrogenase confers resistance to boscalid in Sclerotinia sclerotiorum. Molecular Plant Pathology, 16, 653-661.

Wiemann P, Brown D, Kleigrewe K, Bok J, Keller N, Humpf H U, Tudzynski B. 2010. FfVEL1 and FfLAE1, components of a velvet-like complex in Fusarium fujikuroi, affect differentiation, secondary metabolism and virulence. Molecular microbiology, 77, 972-994.

Wu D, Oide S, Zhang N, Choi M Y, Turgeon, B G. 2012. ChLAE1 and ChVEL1 regulate T-toxin production, virulence, oxidative stress response, and development of the maize pathogen Cochliobolus heterostrophus. PLoS Pathogens, 8, e1002542.

Xu W, Wang K, Wang H, Liu Z, Shi Y, Gao Z, Wang Z G. 2020. Evaluation of the biocontrol potential of Bacillus sp. WB against Fusarium oxysporum f. sp. niveum. Biological Control, 147, 104288.

Yao X, Mi D D , Li Z J, Yang B Y , Zheng Y, Qi Y J, Guo J H. 2019. Comparative Transcriptome Analysis Reveals the Biocontrol Mechanism of Bacillus velezensis F21 Against Fusarium Wilt on Watermelon. Frontiers in Microbiology, 10, 652.

Yu M, Yu J, Cao H, Pan X, Song T, Qi Z Q, Du Y, Huang S W, Liu Y F. 2022. The Velvet Protein UvVea Regulates Conidiation and Chlamydospore Formation in Ustilaginoidea virens. Journal of Fungi, 8, 479.

Zhang R, Xu Q, Zhang Y, Zhu F. 2018. Baseline Sensitivity and Toxic Actions of Prochloraz to Sclerotinia sclerotiorum. Plant Disease, 102, 2149-2157.