中国农业科学 ›› 2019, Vol. 52 ›› Issue (24): 4453-4469.doi: 10.3864/j.issn.0578-1752.2019.24.002
李剑峰1,张博1,全建章2,王永芳2,张小梅1,赵渊1,袁玺垒1,贾小平1(),董志平2(
)
收稿日期:
2019-05-28
接受日期:
2019-06-25
出版日期:
2019-12-16
发布日期:
2020-01-15
通讯作者:
贾小平,董志平
作者简介:
李剑峰,E-mail:1299569468@qq.com。
基金资助:
JianFeng LI1,Bo ZHANG1,JianZhang QUAN2,YongFang WANG2,XiaoMei ZHANG1,Yuan ZHAO1,XiLei YUAN1,XiaoPing JIA1(),ZhiPing DONG2(
)
Received:
2019-05-28
Accepted:
2019-06-25
Online:
2019-12-16
Published:
2020-01-15
Contact:
XiaoPing JIA,ZhiPing DONG
摘要:
【目的】通过在海南省乐东县、河南省洛阳市、吉林省吉林市和公主岭市4个不同地理环境调查谷子10个主要性状,进行SSR标记与性状的关联分析,获得单一环境特异表达位点及多个环境共同表达位点,发掘优异等位变异,探究谷子可能的生态适应性形成机制,为开展谷子分子辅助选择育种奠定基础。【方法】连续2年在吉林省吉林市和公主岭市、河南省洛阳市、海南省乐东县调查102份谷子资源株高、穗长、叶片数、穗粗、抽穗期、穗码数、码粒数、穗重、穗粒重、千粒重10个主要农艺性状的基础上,首先利用SPSS19.0软件对每个地理环境进行性状间相关性分析,再用70个多态性SSR标记对谷子资源进行基因型鉴定,分析102份谷子资源的遗传多样性和群体遗传结构,最后用TASSEL5.0软件进行标记间的连锁不平衡分析,并用GLM和MLM 2种模型开展分子标记与表型性状的关联分析。【结果】除了千粒重,其余9个性状间在4个地理环境普遍存在显著或极显著正相关,千粒重只在在吉林市、公主岭市、洛阳市3个环境与穗粒重、株高、穗长、穗重呈显著或极显著正相关,在海南乐东县与其他9个性状没有显著相关性;70对SSR引物共检测到397个等位基因,平均每个标记的观测等位基因数、有效等位基因数、期望杂合度、Shannon指数分别为6、2.24、0.4637、0.7738;遗传多样性和群体结构分析均将102份谷子材料分为4个群,来自河南的谷子材料散布在4个群中,表现较丰富的遗传多样性;70对SSR标记间连锁不平衡分析未发现明显的连锁不平衡结构;GLM、MLM 2种模型共检测到10个关联标记,结合等位效应分析确定b115、MPGC13、b227、b194、p56在吉林市、公主岭市2个环境与穗重、穗长、叶片数、抽穗期4个性状关联,sigms9034、b125在洛阳市与穗长、穗码数关联,P18和p59在乐东县与穗重关联,p6在吉林市、公主岭市和洛阳市分别与叶片数和穗码数关联,单个标记对表型变异的平均贡献率在7.76%—34.05%;3个标记等位基因sigms9034-168、P18-166、p56-244分别能够显著增加穗长、穗重,缩短抽穗期。【结论】连续2年获得了5个在吉林市、公主岭市稳定表达的标记位点b115、MPGC13、b227、b194和p56,2个在洛阳市稳定表达的标记位点sigms9034、b125,2个在乐东县稳定表达的位点P18和p59,1个在洛阳市和吉林市、公主岭市同时稳定表达的标记位点p6;获得了3个可用于穗部性状、生育期标记辅助选择的优异等位基因sigms9034(168 bp)、P18(166 bp)、p56(244 bp)。
李剑峰,张博,全建章,王永芳,张小梅,赵渊,袁玺垒,贾小平,董志平. 基于SSR标记的谷子主要农艺性状关联位点检测及等位变异分析[J]. 中国农业科学, 2019, 52(24): 4453-4469.
JianFeng LI,Bo ZHANG,JianZhang QUAN,YongFang WANG,XiaoMei ZHANG,Yuan ZHAO,XiLei YUAN,XiaoPing JIA,ZhiPing DONG. Associated Loci Detection and Elite Allelic Variations Analysis of Main Agronomic Traits in Foxtail Millet (Setaria italica L.) Based on SSR Markers[J]. Scientia Agricultura Sinica, 2019, 52(24): 4453-4469.
附表1
102份谷子材料表Attached table 1 102 cultivars of foxtail millet"
品种编号 Variety number | 品种名 Variety name | 来源地 Origin area | 品种编号 Variety number | 品种名 Variety name | 来源地 Origin area |
---|---|---|---|---|---|
HN1 | 郑05-1 Zheng05-1 | 中国河南 Henan, China | HB21 | 104 104 | 未知 Unknown |
HN2 | 郑05-2 Zheng05-2 | 中国河南 Henan, China | HB22 | 200152 200152 | 未知 Unknown |
HN3 | 安04-4705 An04-4705 | 中国河南 Henan, China | HB23 | 8322-14 8322-14 | 未知 Unknown |
HN4 | 安04-4852 An04-4852 | 中国河南 Henan, China | HB24 | 冀谷32 Jigu32 | 中国河北 Hebei, China |
HN5 | 安04-5014 An04-5014 | 中国河南 Henan, China | HB25 | 冀谷30 Jigu30 | 中国河北 Hebei, China |
HN6 | 豫谷1 Yugu1 | 中国河南 Henan, China | HB26 | 冀谷28 Jigu28 | 中国河北 Hebei, China |
HN7 | 安09-8525 An09-8525 | 中国河南 Henan, China | HB27 | 冀谷27 Jigu27 | 中国河北 Hebei, China |
HN8 | 安08-4125 An08-4125 | 中国河南 Henan, China | HB28 | 95307 95307 | 中国河北 Hebei, China |
HN9 | 郑07-2 Zheng07-2 | 中国河南 Henan, China | HB29 | 谷丰2 Gufeng2 | 中国河北 Hebei, China |
HN10 | 郑谷2 Zhenggu2 | 中国河南 Henan, China | HB30 | K325 K325 | 未知 Unknown |
HN11 | 豫谷6 Yugu6 | 中国河南 Henan, China | HB31 | 白米1号 Baimi 1 hao | 中国河北 Hebei, China |
HN12 | 安10-4240 An10-4240 | 中国河南 Henan, China | HB32 | 冀谷1 Jigu1 | 中国河北 Hebei, China |
HN13 | 郑07-1 Zheng07-1 | 中国河南 Henan, China | HB33 | 冀特5 Jite5 | 中国河北 Hebei, China |
HN14 | 安10-4172 An10-4172 | 中国河南 Henan, China | HB34 | 坝谷139 Bagu139 | 中国河北 Hebei, China |
HN15 | 豫谷17 Yugu17 | 中国河南 Henan, China | HB35 | 坝91-0079 Ba91-0079 | 中国河北 Hebei, China |
HN16 | 豫谷11 Yugu11 | 中国河南 Henan, China | HB36 | 毛谷2 Maogu2 | 中国河北 Hebei, China |
HN17 | 豫谷8 Yugu8 | 中国河南 Henan, China | HB37 | 大白谷 Dabaigu | 中国河北 Hebei, China |
HN18 | 郑06-6 Zheng06-6 | 中国河南 Henan, China | HB38 | 大九根齐 Dajiugenqi | 中国河北 Hebei, China |
HN19 | 豫谷3 Yugu3 | 中国河南 Henan, China | HB39 | 坝谷6 Bagu6 | 中国河北 Hebei, China |
HN20 | 安4117 An4117 | 中国河南 Henan, China | HB40 | 四留钱 Siliuqian | 中国河北 Hebei, China |
HN21 | 安-4585 An-4585 | 中国河南 Henan, China | HB41 | 坝谷210 Bagu210 | 中国河北 Hebei, China |
HN22 | 安5424 An5424 | 中国河南 Henan, China | SD1 | 鲁谷10 Lugu10 | 中国山东 Shandong, China |
HN23 | 豫谷15 Yugu15 | 中国河南 Henan, China | SD2 | 早白糯 Zaobainuo | 中国山东 Shandong, China |
HN24 | 豫谷13 Yugu13 | 中国河南 Henan, China | SD3 | 济大粒2 Jidali2 | 中国山东 Shandong, China |
HN25 | 豫谷16 Yugu16 | 中国河南 Henan, China | SD4 | 济叶冲4 Jiyechong4 | 中国山东 Shandong, China |
HN26 | 豫谷18 Yugu18 | 中国河南 Henan, China | SD5 | 济叶冲20 Jiyechong20 | 中国山东 Shandong, China |
HN27 | 郑州12 Zhengzhou12 | 中国河南 Henan, China | SD6 | 济长穗8 Jichangsui8 | 中国山东 Shandong, China |
HN28 | 郑315 Zheng315 | 中国河南 Henan, China | SD7 | 济丰30 Jifeng30 | 中国山东 Shandong, China |
HN29 | 郑8041 Zheng8041 | 中国河南 Henan, China | SD8 | 金线子 Jinxianzi | 中国山东 Shandong, China |
HN30 | 豫谷5 Yugu5 | 中国河南 Henan, China | SD9 | 鲁谷3 Lugu3 | 中国山东 Shandong, China |
HN31 | 新农673 Xinnong673 | 中国河南 Henan, China | SD10 | 鲁谷7 Lugu7 | 中国山东 Shandong, China |
HB1 | 谷丰1 Gufeng1 | 中国河北 Hebei, China | SD11 | 书香1号 Shuxiang1hao | 中国山东 Shandong, China |
HB2 | 71杂30-2 71za30-2 | 未知 Unknown | SC1 | 11郄961 11 xi 961 | 未知 Unknown |
HB3 | 张庙谷选变Zhangmiaoguxuanbian | 未知 Unknown | SC2 | 11郄1071 11 xi 1071 | 未知 Unknown |
HB4 | 冀谷15 Jigu15 | 中国河北 Hebei, China | BJ1 | 06-766 06-766 | 中国北京 Beijing, China |
HB5 | 冀谷17 Jigu17 | 中国河北 Hebei, China | BJ2 | 小早谷 Xiaozaogu | 中国北京 Beijing, China |
HB6 | 衡谷9 Jigu9 | 中国河北 Hebei, China | NMG1 | 小香米 Xiaoxiangmi | 中国河北 Hebei, China |
HB7 | 金谷1 Jingu1 | 中国河北 Hebei, China | NMG2 | 籼紫灰谷Shanzihuigu | 中国内蒙古 Neimenggu, China |
HB8 | 沧372 Cang372 | 中国河北 Hebei, China | NMG3 | 二白谷 Erbaigu | 中国内蒙古 Neimenggu, China |
HB9 | 沧369 Cang369 | 中国河北 Hebei, China | NMG4 | 黄玉3 Huangyu3 | 中国内蒙古 Neimenggu, China |
HB10 | 冀谷29 Jigu29 | 中国河北 Hebei, China | SX1 | 呼和浩特大毛谷Huhehaotedamaogu | 中国陕西 Shanxi, China |
HB11 | 冀谷26 Jigu26 | 中国河北 Hebei, China | SX2 | 延谷4 Yangu4 | 中国陕西 Shanxi, China |
HB12 | 冀谷24 Jigu24 | 中国河北 Hebei, China | SX3 | 红杆谷 Honggangu | 中国陕西 Shanxi, China |
HB13 | 冀谷22 Jigu22 | 中国河北 Hebei, China | MFGY1 | ISE430 ISE430 | 美国 America |
HB14 | 冀谷19 Jigu19 | 中国河北 Hebei, China | MFGY2 | 法谷28-81 Fagu28-81 | 法国 French |
HB15 | 冀谷18 Jigu18 | 中国河北 Hebei, China | MFGY3 | SET3/80 SET3/80 | 德国 Germany |
HB16 | 冀谷31 Jigu31 | 中国河北 Hebei, China | MFGY4 | ISE775 ISE775 | 印度 India |
HB17 | 复12 Fu12 | 中国河北 Hebei, China | HCCR1 | 龙谷26 Longgu26 | 中国黑龙江 Heilongjiang, China |
HB18 | 冀创1 Jichuang1 | 中国河北 Hebei, China | HCCR2 | 朝鲜谷子 Chaoxianguzi | 朝鲜 Korea |
HB19 | 2015 2015 | 中国河北 Hebei, China | HCCR3 | 谷绿早1 Gulvzao1 | 朝鲜 Korea |
HB20 | 2013 2013 | 中国河北 Hebei, China | HCCR4 | 大王国 Dawangguo | 日本 Japan |
附表2
70对SSR引物Attached table 2 70 pairs of SSR primers"
SSR标记 SSR markers | 引物序列 Primer sequence (5’-3’) | SSR标记 SSR markers | 引物序列 Primer sequence (5’-3’) |
---|---|---|---|
b115 | F:GGTAGCGACGGATCTACAGC R:GCTAGCAAATGCTGTCATGG | b202 | F:AGAGCCCACGTCAAACC R:AAACTGGACTAGAAGAAGCATAG |
b116 | F:GCAAGCGTGATGTCAGATTAT R:ATAGGATGGTGGAAGCCCA | b101 | F:ATCTAGGTGCCGATGCGT R:TGTGGGAAGAAGCTAGGGAA |
b122 | F:ACTTCTTCCTTCCTTGCGG R:TGTGGGATTAAGGTGCATCG | b106 | F:TGCTTTGCTCTCTTCTCTCA R:ACGGACGATGAGGAATTGT |
b125 | F:GCCATGAAACAGGTACAAAAGG R:GCATCCCCTTAATTTGTCAATG | p59 | F:TAATTTTGTGGCGTGGGATG R:GCACTGGTTTTGTTGAATGG |
p6 | F:AAGGATGGAATTTGCCACTG R:TTTCGACGATTTGCTTCAAC | p98 | F:ATTCATCAGTAGCACAGC R:TGGAACTAAGAACAGGAAAC |
p5 | F:CTTCCCTCCCTCCCTGAC R:CTGAGCTGAGCTGCCTTTG | p56 | F:GATGTGTACGGGTTGCATTG R:TGGGTTTCAGGGCTCTCTC |
p3 | F:GCAGAAAGCATGCCGTAGTC R:GCTTGGAGTCCACATGGATAG | p61 | F:CATCCGCGTCATCTGAATC R:ACCTGCTGCTATCCATCACC |
p8 | F:CGATCGAATGATCGATGAAC R:CCCTTTGTCCGATCACGTC | p68 | F:CATGCGTTGATCGTTTGTG R:ACCACGCATTTACATGATCG |
p10 | F:CAATCACATCCGAGCATTTC R:CACCCACCGTGTTGATCTG | p74 | F:CAACCTAGTTTGCCTCAGTTATTC R:AGTCACGTACATGGGTGCAG |
p12x | F:ACGAGTCACAAATCACAGCAC R:ATGCCTGAGCGGAACGGAA | p80 | F:GCCGTTGGATTTGATTATGG R:TGTGGTTAGTTTATGTGGCTTG |
p16 | F:TTTCTCCCTCTCTCGATTCC R:AAATTGGCGTGCTAACAACC | p87 | F:ACCTTTGACAAACGAGACACG R:GTTCGACTTGCATTGACTGG |
b127 | F:CCTCAAGTCAGTGAGATGCAA R:CAGAGCTGTTTAATCCTTGTTCA | p89 | F:GCCTGTCTGAAAATTCTCAATG R:AGACGTGACATTAGCGCTTG |
b129 | F:CACACTCTTCTCCCCTTTTCC R:ACGGTAACGGAGGATGGCTA | p88 | F:CAAGCCACCCAGTCTAGAGG R:TTCATCAGAACTGCGCAAAC |
b142 | F:TGGTAAAACTCCCATATTGAGC R:GCCCCATCCTTGATAACAGA | p92 | F:TGGAATTGGAACCCTTTCG R:GCCATGCAAACAGTACCATC |
b147 | F:CTACTGCCTTCTGGCCTCC R:GGGCATTCTTGCTTCAGTCA | p85 | F:GAATTAGGCCGATGCACAAC R:ATCCTAACTGCATGGCAAGG |
b158 | F:GATGAGGAAAAGGTAGGTTGGA R:CTGCAACGTGCAGAACTACG | p100 | F:AGTTGACACCACACATAACAA R:AGAATACTCCTACCTGCCAC |
b161 | F:GGCATAAAAGTAAAAACCAACCA R:ACCTGGCTTCTGTCAGTGAA | p95X | F:GTCTCTGATAGTGCTTGAGCG R:ACAGGGATGAAGGGCGATG |
b163 | F:CTCGGAAGCTCAGATTCTCC R:CACTTCCTGCAGCTCTCACA | p29 | F:GATGAGCACACGTTGATTGG R:GGACTTCACCACCGAGATG |
b165 | F:GCTTTGGTTTGGTTTGGTTGG R:CCATTAGTCTCTGCCCTTGTT | p44 | F:TTCCCGGAACAGACAAGAAC R:GCGTTGGAAGCCATGGAG |
b166 | F:CGCCCATACTACCCAACAG R:ACCTCACCTTCCACTCCTC | p42 | F:GCGACTTTCCCCTTCCAATC R:TTCCTTTTGTTGGCTTCTCC |
b194 | F:CTGGGTTCCGTCTACCGTA R:CACACCCGAAGAGGCAAAG | p91 | F:AGCTGTGCTCCTCTGATCTTG R:TAACGTGGGGATGCACTAGC |
b200 | F:CATCGATCTCAACCTGTCCTT R:ATGAGCCGTCATGTCACAAA | sigms11655 | F:TCGTTTAAGCTGGAATTGGG R:AGAGTACCGTCGGCGTCTAA |
b107 | F:AGAACGAGGTGGTGTGTGG R:GGGTCTCACGCTCTCATCA | P18 | F:TTCTCTCGTTGGAATTTTTGTG R:GGAACAGATATCCTTTTCACTCTT |
b123 | F:GGTGTTCTCCTGTGTGC R:AGAGTTATTTCCAGCATTAGTG | p20 | F:GTGCCCGCTTAGCTTTAATC R:ATGCACGTGGGACCCATAC |
b159 | F:GCCAGTCCGAGATGGTTAAG R:AGCTCTAGCAGTTGGGGACA | p17x | F:CGGACACCTGAAAGACGAA R:GTCACTTGTTGTTGTTGCG |
b190 | F:GAAATTTCACAAGTGTTGGTG R:TGATCGGAGCAGAGTGTTGA | sigms9034 | F:TGCTGGTCGCAGTACTTGAT R:TCCTCTGCTCTGCTCTCCTC |
b234 | F:GCCGCAACGAACAACCG R:CCTGTCCCTATCCCTGTCG | sigms10291 | F:TCGTCTCCGTCAGTAATCCC R:GCAGCAGAAGCAGATGAGG |
b255 | F:GAGGACAGCGGCCATT R:CCTCCCTCCATTTACTTTGG | sigms11641 | F:CACCATGGCCACCATATGTA R:TGGTTGCTCAGCATAGCCTT |
b242 | F:CACTACCACTGTTCCAGATCG R:CAGGGACCTTGCTTGCATAC | sigms11693 | F:CTAATTTTGCATGTGCCAGG R:CCACATACACCATGAGGTTCC |
b185 | F:GCACGTGTGACTTTCCACAT R:GTGAATGGCACACGAAACTG | sigms11672 | F:TAGTAAACTTGGGCCCATCC R:ATAACTTCTCCCCCACCACC |
b227 | F:TGATCTGGCAGAACGAACA R:CAATTCCTGGACCAATATGC | sigms10287 | F:GACCGATTCCTTCCAAACAA R:TGTCAGCTGGAGATGGTGTC |
b233 | F:GCCACGCACACCAACTT R:CTCCCGCAGAACACGCA | MPGD46 | F:GATGGGTCGTCGTTAGAGTTT R:GGAAAGGGAAAGGAAGATAGC |
b258 | F:GGGCCAATAATGGTTGCATA R:TTGCACATCCAAATCTTTCC | MPGD10 | F:CATCTGTTGCTGCTGATGCT R:AATTCTCAAAAGAGAGACCCACTG |
b153 | F:ACCCAACACATTCTCCTGAA R:TGCTATCAAAATAGTGCTAGAAT | sigms11 | F:AGATCCTCGCAGACTTCGTC R:TGCAATCACCAGAGAGAAGC |
b187 | F:TTGGACAAATGACGCTATGC R:CTGCATCAAATCAGGACCAC | MPGC13 | F:CTGAAGCATTCGATAAACTAGCTG R:GTAGTGAAAAATGAGCAAGGGTCT |
表1
不同地理环境谷子主要农艺性状相关性分析"
环境 Environments | 性状Trait | HS | GW | PH | NL | PL | PD | SN | GN | SW | 1000-GW |
---|---|---|---|---|---|---|---|---|---|---|---|
吉林市 Jilin | HS | 1 | |||||||||
GW | -0.157 | 1 | |||||||||
PH | 0.035 | 0.340** | 1 | ||||||||
NL | 0.538** | 0.195 | 0.105 | 1 | |||||||
PL | 0.04 | 0.464** | 0.530** | 0.109 | 1 | ||||||
PD | 0.173 | 0.415** | 0.012 | 0.366** | 0.208 | 1 | |||||
SN | 0.421** | 0.101 | 0.116 | 0.299* | 0.285* | 0.033 | 1 | ||||
GN | -0.08 | 0.667** | 0.129 | 0.283* | 0.119 | 0.547** | -0.209 | 1 | |||
SW | -0.09 | 0.957** | 0.361** | 0.254* | 0.547** | 0.478** | 0.122 | 0.694** | 1 | ||
1000-GW | -0.315** | 0.388** | 0.242* | -0.226 | 0.321** | 0.051 | -0.269* | -0.005 | 0.382** | 1 | |
公主岭市 Gongzhuling | HS | 1 | |||||||||
GW | 0.367** | 1 | |||||||||
PH | 0.420** | 0.343** | 1 | ||||||||
NL | 0.756** | 0.552** | 0.485** | 1 | |||||||
PL | 0.329* | 0.332** | 0.686** | 0.282* | 1 | ||||||
PD | 0.571** | 0.480** | 0.173 | 0.520** | 0.260* | 1 | |||||
SN | 0.363** | 0.206 | 0.380** | 0.342** | 0.302* | -0.115 | 1 | ||||
GN | 0.323* | 0.657** | 0.064 | 0.401** | 0.077 | 0.594** | -0.170 | 1 | |||
SW | 0.437** | 0.938** | 0.373** | 0.596** | 0.395** | 0.618** | 0.132 | 0.674** | 1 | ||
1000-GW | -0.018 | 0.307* | 0.383** | 0.068 | 0.427** | 0.092 | 0.202 | 0.012 | 0.384** | 1 | |
洛阳市 Luoyang | HS | 1 | |||||||||
GW | 0.552** | 1 | |||||||||
PH | 0.571** | 0.575** | 1 | ||||||||
NL | 0.662** | 0.742** | 0.670** | 1 | |||||||
PL | 0.338** | 0.205 | 0.716** | 0.333** | 1 | ||||||
PD | 0.567** | 0.688** | 0.558** | 0.761** | 0.388** | 1 | |||||
SN | 0.565** | 0.554** | 0.667** | 0.648** | 0.513** | 0.541** | 1 | ||||
GN | 0.281* | 0.745** | 0.302** | 0.493** | -0.079 | 0.544** | 0.180 | 1 | |||
SW | 0.619** | 0.951** | 0.672** | 0.794** | 0.363** | 0.766** | 0.614** | 0.687** | 1 | ||
1000-GW | 0.211 | 0.347** | 0.343** | 0.300** | 0.330** | 0.296** | 0.264* | 0.046 | 0.371** | 1 | |
乐东县Ledong | HS | 1 | |||||||||
GW | 0.623** | 1 | |||||||||
PH | 0.163 | 0.228* | 1 | ||||||||
NL | 0.771** | 0.781** | 0.230* | 1 | |||||||
PL | 0.607** | 0.754** | 0.189 | 0.609** | 1 | ||||||
PD | 0.665** | 0.858** | 0.244** | 0.845** | 0.613** | 1 | |||||
SN | 0.429** | 0.553** | 0.284* | 0.534** | 0.558** | 0.442** | 1 | ||||
GN | 0.353** | 0.735** | 0.159 | 0.557** | 0.364** | 0.697** | 0.132 | 1 | |||
SW | 0.622** | 0.808** | 0.292* | 0.708** | 0.672** | 0.775** | 0.557** | 0.503** | 1 | ||
1000-GW | -0.003 | 0.194 | -0.083 | 0.064 | 0.120 | 0.129 | 0.079 | -0.008 | 0.122 | 1 |
表2
70对SSR标记在102份谷子材料中的多态性"
SSR标记 SSR markers | 等位基因数 Allele number | 有效等位基因数 Number of effective alleles | Shannon指数 Shannon index | 期望杂合度 Expected heterozygosity |
---|---|---|---|---|
b115 | 6 | 2.005 | 0.618 | 0.350 |
b116 | 4 | 1.712 | 0.521 | 0.325 |
b122 | 5 | 2.200 | 0.719 | 0.431 |
b125 | 5 | 2.447 | 0.926 | 0.575 |
p6 | 4 | 2.059 | 0.738 | 0.468 |
p5 | 6 | 2.022 | 0.632 | 0.363 |
p3 | 8 | 2.776 | 1.026 | 0.602 |
p8 | 6 | 1.983 | 0.611 | 0.346 |
p10 | 5 | 2.204 | 0.786 | 0.494 |
p12x | 8 | 2.226 | 0.767 | 0.448 |
p16 | 5 | 1.849 | 0.631 | 0.390 |
b127 | 7 | 2.355 | 0.765 | 0.438 |
b129 | 4 | 2.283 | 0.849 | 0.516 |
b142 | 7 | 2.595 | 0.903 | 0.522 |
b147 | 4 | 1.773 | 0.622 | 0.400 |
b158 | 5 | 2.148 | 0.784 | 0.495 |
b161 | 5 | 1.926 | 0.666 | 0.405 |
b163 | 4 | 1.909 | 0.568 | 0.332 |
b165 | 4 | 1.827 | 0.590 | 0.395 |
b166 | 4 | 1.943 | 0.685 | 0.442 |
b194 | 6 | 1.909 | 0.657 | 0.380 |
b200 | 6 | 2.277 | 0.802 | 0.470 |
b107 | 7 | 1.961 | 0.721 | 0.446 |
b123 | 6 | 2.426 | 0.872 | 0.520 |
b159 | 6 | 2.600 | 0.900 | 0.509 |
b190 | 5 | 1.933 | 0.696 | 0.433 |
b234 | 4 | 1.742 | 0.503 | 0.308 |
b255 | 6 | 2.434 | 0.816 | 0.469 |
b242 | 4 | 2.100 | 0.766 | 0.477 |
b185 | 4 | 2.153 | 0.803 | 0.512 |
b227 | 4 | 2.025 | 0.637 | 0.392 |
b233 | 5 | 2.354 | 0.895 | 0.558 |
b258 | 6 | 2.118 | 0.775 | 0.478 |
b153 | 5 | 2.497 | 0.854 | 0.498 |
b187 | 5 | 2.498 | 0.940 | 0.578 |
b202 | 7 | 2.329 | 0.790 | 0.449 |
b101 | 6 | 2.218 | 0.779 | 0.448 |
b106 | 8 | 2.325 | 0.775 | 0.448 |
p59 | 5 | 2.258 | 0.812 | 0.496 |
p98 | 4 | 2.177 | 0.728 | 0.443 |
p56 | 5 | 2.123 | 0.806 | 0.508 |
p61 | 4 | 1.671 | 0.498 | 0.319 |
p68 | 5 | 2.098 | 0.734 | 0.466 |
p74 | 5 | 2.366 | 0.825 | 0.515 |
p80 | 8 | 2.797 | 1.025 | 0.598 |
p87 | 8 | 3.092 | 1.118 | 0.642 |
p89 | 8 | 2.538 | 0.892 | 0.497 |
p88 | 8 | 2.832 | 0.974 | 0.560 |
p92 | 7 | 2.425 | 0.843 | 0.498 |
p85 | 6 | 2.286 | 0.838 | 0.503 |
p100 | 6 | 2.448 | 0.879 | 0.530 |
p95X | 9 | 2.827 | 0.935 | 0.502 |
p29 | 7 | 2.714 | 0.964 | 0.534 |
p44 | 8 | 2.932 | 1.036 | 0.605 |
p42 | 7 | 2.785 | 0.989 | 0.567 |
p91 | 8 | 2.362 | 0.802 | 0.474 |
sigms11655 | 6 | 1.788 | 0.490 | 0.295 |
P18 | 3 | 1.519 | 0.439 | 0.279 |
p20 | 7 | 2.519 | 0.937 | 0.570 |
p17x | 6 | 2.335 | 0.818 | 0.495 |
sigms9034 | 5 | 1.593 | 0.493 | 0.311 |
sigms10291 | 4 | 2.174 | 0.728 | 0.451 |
sigms11641 | 3 | 1.726 | 0.538 | 0.368 |
sigms11693 | 6 | 2.463 | 0.946 | 0.572 |
sigms11672 | 4 | 1.686 | 0.516 | 0.321 |
sigms10287 | 7 | 2.723 | 0.913 | 0.498 |
MPGD46 | 6 | 2.297 | 0.804 | 0.466 |
MPGD10 | 5 | 2.649 | 0.974 | 0.588 |
sigms11 | 6 | 2.390 | 0.883 | 0.522 |
MPGC13 | 5 | 1.794 | 0.599 | 0.355 |
表3
4个地理环境GLM、MLM模型关联分析结果"
性状 Traits | 关联标记 Associated markers | 染色体 Chromosome | 显著性检验值(P) Significance test value (P) | 表型贡献率(R2) Phenotypic variation (R2) | 环境 Environments | 年份 Years | 模型 Model |
---|---|---|---|---|---|---|---|
GW SW PL | b115 | 2 | 9.99E-04 0.021 9.99E-04 0.013 0.0317 1.54E-04 | 0.4355 0.2456 0.4661 0.2521 0.0807 0.2421 | 吉林Jilin 公主岭Gongzhuling 吉林Jilin 公主岭Gongzhuling 吉林Jilin 公主岭Gongzhuling | 2015 2016 2015 2016 2015 2016 | GLM MLM |
NL | MPGC13 | 7 | 9.99E-04 9.99E-04 | 0.24 0.3156 | 吉林Jilin 公主岭Gongzhuling | 2015 2016 | GLM |
PL PH NL SN | p6 | 8 | 9.99E-04 0.025 0.0057 0.0118 0.0123 8.46E-04 0.033 0.0171 0.0127 0.015 | 0.1958 0.1712 0.0718 0.1109 0.0701 0.1591 0.0778 0.0774 0.0943 0.1031 | 吉林Jilin 公主岭Gongzhuling 吉林Jilin 公主岭Gongzhuling 吉林Jilin 公主岭Gongzhuling 吉林Jilin 公主岭Gongzhuling 洛阳Luoyang 洛阳Luoyang | 2015 2016 2015 2016 2015 2016 2015 2016 2015 2016 | GLM MLM |
PL | sigms9034 | 6 | 9.99E-04 0.009 | 0.2361 0.1898 | 洛阳Luoyang 洛阳Luoyang | 2015 2016 | GLM |
PL SN | b125 | 5 | 0.026 0.014 0.0094 0.0073 | 0.2717 0.2409 0.1405 0.1645 | 洛阳Luoyang 洛阳Luoyang 洛阳Luoyang 洛阳Luoyang | 2015 2016 2015 2016 | GLM MLM |
HS | b227 | 1 | 9.99E-04 9.99E-04 0.0372 0.0308 | 0.2460 0.3075 0.0999 0.0811 | 吉林Jilin 公主岭Gongzhuling 吉林Jilin 公主岭Gongzhuling | 2015 2016 2015 2016 | GLM MLM |
HS | b194 | 9 | 0.01 0.032 | 0.1537 0.0874 | 吉林Jilin 公主岭Gongzhuling | 2015 2016 | MLM |
HS | p56 | 2 | 0.0215 0.0204 | 0.1202 0.0936 | 吉林Jilin 公主岭Gongzhuling | 2015 2016 | MLM |
SW | P18 | 5 | 0.0117 0.0039 | 0.0861 0.1044 | 乐东Ledong 乐东Ledong | 2015 2016 | MLM |
SW | p59 | 未知 Unknown | 0.0452 0.016 | 0.0894 0.1171 | 乐东Ledong 乐东Ledong | 2015 2016 | MLM |
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