中国农业科学 ›› 2019, Vol. 52 ›› Issue (22): 4027-4038.doi: 10.3864/j.issn.0578-1752.2019.22.008
收稿日期:
2019-06-05
接受日期:
2019-07-08
出版日期:
2019-11-16
发布日期:
2019-11-16
通讯作者:
管延安
作者简介:
秦岭,0531-66659029;E-mail:qinling1021@163.com
基金资助:
QIN Ling,ZHANG YanTing,CHEN ErYing,YANG YanBing,LI FeiFei,GUAN YanAn()
Received:
2019-06-05
Accepted:
2019-07-08
Online:
2019-11-16
Published:
2019-11-16
Contact:
YanAn GUAN
摘要:
【目的】确定谷子萌发期耐盐评价指标,筛选萌发期耐盐种质,并探讨不同基因型谷子苗期盐胁迫对保护酶系统的影响,为谷子大规模耐盐性鉴定、耐盐机理的研究提供鉴定方法和优异资源。【方法】 以不同生态区的54份谷子种质为试验材料,用1.5%NaCl溶液进行盐胁迫,蒸馏水为对照,采用培养皿发芽法在人工气候培养箱内进行谷子萌发期耐盐性鉴定;测定谷子相对发芽势、相对发芽率、相对胚芽长、相对胚根长、相对胚芽比以及发芽率、盐害率等指标;通过对指标值的相关性分析、主成分分析和聚类分析,筛选谷子萌发耐盐评价指标。采用筛选出的3个不同耐盐性谷子品种,以0.5%NaCl溶液进行苗期盐胁迫,测定盐土盆栽条件下苗期叶片超氧化物歧化酶(SOD)、过氧化物酶(POD)、过氧化氢酶(CAT)活性。分析盐胁迫下3个基因型间生理响应机理的差异。【结果】 在1.5%NaCl溶液胁迫下,谷子的相对发芽势与相对发芽率(r=0.51,P<0.01)、相对胚芽长之间呈显著正相关(r=0.54,P<0.01);相对胚芽长与相对胚根长呈显著正相关(r=0.64,P<0.01);发芽率盐害率与相对发芽势(r=-0.37,P<0.01)、相对胚芽长(r=-0.51,P<0.01)呈显著负相关。通过主成分分析将萌发期盐胁迫处理的6个单项指标转换成3个彼此独立的综合指标;通过隶属函数分析,得到不同品种萌发期耐盐性综合评价值(D值),并通过聚类分析,将54份谷子品种分成高度盐敏感品种、盐敏感品种、中度耐盐品种、耐盐品种以及高度耐盐品种5个不同类型。其中,高度耐盐品种有4个,分别是华北夏谷区的济谷16、矮88,西北春谷区的陇谷3号和延谷13。苗期盐土盆栽试验表明,盐胁迫条件下谷子叶片SOD、POD、CAT酶活性呈现先上升后下降的趋势。耐盐性强品种济谷16的SOD、POD、CAT酶活性上升幅度显著大于耐盐性弱的品种鲁谷1号。【结论】 54份谷子种质材料在耐盐性上存在显著差异,利用隶属函数法综合分析萌发期各指标,全面地评价谷子种质资源萌发期的耐盐性。不同基因型谷子品种叶片保护酶系统对NaCl胁迫响应能力的差异,可能是由于谷子耐盐能力不同造成的。
秦岭, 张艳亭, 陈二影, 杨延兵, 黎飞飞, 管延安. 谷子萌发期耐盐种质筛选及其保护酶系统对盐胁迫的响应[J]. 中国农业科学, 2019, 52(22): 4027-4038.
QIN Ling, ZHANG YanTing, CHEN ErYing, YANG YanBing, LI FeiFei, GUAN YanAn. Screening for Germplasms Tolerant to Salt at Germination Stage and Response of Protective Enzymes to Salt Stress in Foxtail Millet[J]. Scientia Agricultura Sinica, 2019, 52(22): 4027-4038.
表1
萌发期盐胁迫下谷子各性状的变化范围及变异系数"
指标 Index | 变化范围 Variation range | 平均值 Mean | 变异系数 Coefficient of variance |
---|---|---|---|
相对发芽势 Relative germination potential(%) | 0.00—74.57 | 14.24 | 100.27 |
相对发芽率 Relative germination rate(%) | 3.81—93.77 | 36.15 | 56.96 |
相对胚芽长 Relative coleoptile length(%) | 8.66—59.48 | 24.45 | 36.33 |
相对胚根长 Relative radical root length(%) | 4.52—33.62 | 9.83 | 47.64 |
相对根芽比 Relative radical root / coleoptiles(%) | 19.83—105.46 | 41.67 | 35.88 |
相对盐害率 Relative salt damage rate(%) | 6.33—96.19 | 63.85 | 32.25 |
表2
盐胁迫下谷子萌发期各性状的相关性分析"
相关系数 Correlation coefficient | 相对发芽势 Relative germination potential (X1) | 相对发芽率 Relative germination rate (X2) | 相对胚芽长 Relative coleoptile length (X3) | 相对胚根长 Relative radical root length (X4) | 相对根芽比 Relative radical root / coleoptiles (X5) | 相对盐害率 Relative salt damage rate (X6) |
---|---|---|---|---|---|---|
X1 | 1 | |||||
X2 | 0.51** | 1 | ||||
X3 | 0.54** | 0.46** | 1 | |||
X4 | 0.05 | 0.10 | 0.64** | 1 | ||
X5 | -0.37** | -0.28** | -0.27** | 0.47** | 1 | |
X6 | -0.51** | -1.00** | -0.46** | -0.10 | 0.28** | 1 |
表3
各综合指标特征值、贡献率及特征向量"
项目 Item | CI1 | CI2 | CI3 |
---|---|---|---|
特征值Eigen value | 2.984 | 1.613 | 0.870 |
贡献率Contributive ratio (%) | 49.73 | 26.89 | 14.50 |
累积贡献率 Cumulative contributive ratio (%) | 49.73 | 76.62 | 91.12 |
特征向量Eigenvector X1 | 0.435 | -0.092 | 0.324 |
特征向量Eigenvector X2 | 0.517 | -0.075 | -0.465 |
特征向量Eigenvector X3 | 0.438 | 0.340 | 0.462 |
特征向量Eigenvector X4 | 0.144 | 0.748 | 0.099 |
特征向量Eigenvector X5 | -0.252 | 0.552 | -0.490 |
特征向量Eigenvector X6 | -0.517 | 0.075 | 0.465 |
表4
各品种(系)的综合指标值、权重以及D值"
序号 Number | CI1 | CI2 | CI3 | μ (X1) | μ (X2) | μ (X3) | D | 综合评价 Comprehensive valuation |
---|---|---|---|---|---|---|---|---|
1 | -2.2796 | -0.5447 | 0.4050 | 0.0955 | 0.1295 | 0.5460 | 0.1772 | 高度盐敏感Extremely salt sensitivity |
2 | 1.3460 | 0.2178 | -0.1585 | 0.4899 | 0.2264 | 0.4308 | 0.4028 | 中度耐盐Moderate salt tolerance |
3 | 1.2316 | 0.2696 | -1.8077 | 0.4775 | 0.2330 | 0.0939 | 0.3443 | 中度耐盐Moderate salt tolerance |
4 | 0.0000 | 0.8496 | -0.0200 | 0.3435 | 0.3067 | 0.4591 | 0.3510 | 中度耐盐Moderate salt tolerance |
5 | -0.0623 | -1.2831 | 0.4707 | 0.3367 | 0.0357 | 0.5594 | 0.2833 | 盐敏感Salt sensitivity |
6 | 2.2398 | 3.5566 | 0.6492 | 0.5872 | 0.6506 | 0.5958 | 0.6073 | 高度耐盐High salt tolerance |
7 | -0.4241 | -0.1059 | -0.6293 | 0.2973 | 0.1853 | 0.3346 | 0.2702 | 盐敏感Salt sensitivity |
8 | -0.5756 | 1.9376 | 2.6273 | 0.2809 | 0.4449 | 1.0000 | 0.4437 | 耐盐Salt tolerance |
9 | -0.3235 | -0.5754 | -0.4567 | 0.3083 | 0.1256 | 0.3699 | 0.2642 | 盐敏感Salt sensitivity |
10 | -1.0946 | 0.0800 | -0.5070 | 0.2244 | 0.2089 | 0.3596 | 0.2413 | 盐敏感Salt sensitivity |
11 | -0.1767 | 0.4078 | -1.9838 | 0.3243 | 0.2505 | 0.0579 | 0.2601 | 盐敏感Salt sensitivity |
12 | 1.5976 | -0.0443 | -1.1515 | 0.5173 | 0.1931 | 0.2279 | 0.3756 | 中度耐盐Moderate salt tolerance |
13 | 0.6795 | -0.2044 | -0.7963 | 0.4174 | 0.1727 | 0.3005 | 0.3266 | 中度耐盐Moderate salt tolerance |
14 | -0.4494 | -1.1064 | 1.4951 | 0.2946 | 0.0581 | 0.7687 | 0.3002 | 盐敏感Salt sensitivity |
15 | 2.0224 | 0.6177 | 0.1566 | 0.5635 | 0.2772 | 0.4952 | 0.4682 | 耐盐Salt tolerance |
16 | 1.7565 | -1.1427 | 0.0451 | 0.5346 | 0.0535 | 0.4724 | 0.3827 | 中度耐盐Moderate salt tolerance |
17 | 0.9306 | -0.6810 | -0.2228 | 0.4447 | 0.1122 | 0.4177 | 0.3423 | 中度耐盐Moderate salt tolerance |
18 | -1.4052 | -0.1881 | 0.5329 | 0.1906 | 0.1748 | 0.5721 | 0.2466 | 盐敏感Salt sensitivity |
19 | -0.7363 | 6.3065 | 0.4175 | 0.2634 | 1.0000 | 0.5485 | 0.5261 | 高度耐盐High salt tolerance |
20 | 0.4163 | -0.8748 | 0.1120 | 0.3888 | 0.0876 | 0.4861 | 0.3154 | 中度耐盐Moderate salt tolerance |
21 | 2.619 | -1.4495 | -2.1504 | 0.6284 | 0.0145 | 0.0238 | 0.3511 | 中度耐盐Moderate salt tolerance |
22 | -0.0984 | -0.9923 | 0.3280 | 0.3328 | 0.0726 | 0.5302 | 0.2874 | 盐敏感Salt sensitivity |
23 | 2.0215 | -0.3289 | 0.3369 | 0.5634 | 0.1569 | 0.5320 | 0.4385 | 耐盐 Salt tolerance |
24 | 0.8321 | -1.5640 | 0.2352 | 0.4340 | 0.0000 | 0.5113 | 0.3182 | 中度耐盐Moderate salt tolerance |
25 | -0.3626 | -0.8436 | -0.2953 | 0.3040 | 0.0915 | 0.4029 | 0.2570 | 盐敏感Salt sensitivity |
26 | -0.6926 | -1.2175 | 0.2139 | 0.2681 | 0.0440 | 0.5069 | 0.2400 | 盐敏感Salt sensitivity |
27 | -1.9623 | -0.6357 | -0.0949 | 0.1300 | 0.1179 | 0.4438 | 0.1764 | 高度盐敏感Extremely salt sensitivity |
28 | 0.4516 | 1.1999 | -0.8049 | 0.3926 | 0.3512 | 0.2987 | 0.3655 | 中度耐盐Moderate salt tolerance |
29 | 1.2783 | -0.4670 | 0.5348 | 0.4826 | 0.1394 | 0.5725 | 0.3956 | 中度耐盐Moderate salt tolerance |
30 | -1.1852 | -0.1785 | 0.5406 | 0.2145 | 0.1760 | 0.5737 | 0.2603 | 盐敏感Salt sensitivity |
31 | -3.1570 | 1.8624 | -2.2671 | 0.0000 | 0.4353 | 0.0000 | 0.1285 | 高度盐敏感Extremely salt sensitivity |
32 | -3.0920 | -0.8026 | -0.0456 | 0.0071 | 0.0967 | 0.4539 | 0.1046 | 高度盐敏感Extremely salt sensitivity |
33 | -1.5469 | -1.1680 | 0.6173 | 0.1752 | 0.0503 | 0.5893 | 0.2042 | 高度盐敏感Extremely salt sensitivity |
34 | -1.8683 | -1.2398 | 1.1237 | 0.1402 | 0.0412 | 0.6928 | 0.1989 | 高度盐敏感Extremely salt sensitivity |
35 | 1.0959 | 1.4691 | -0.4824 | 0.4627 | 0.3854 | 0.3646 | 0.4243 | 耐盐Salt tolerance |
36 | -0.5245 | -0.4524 | 0.4839 | 0.2864 | 0.1412 | 0.5621 | 0.2874 | 盐敏感Salt sensitivity |
37 | -1.9564 | -0.5711 | 0.2578 | 0.1306 | 0.1262 | 0.5159 | 0.1906 | 高度盐敏感Extremely salt sensitivity |
38 | 5.6668 | -1.3401 | 0.1074 | 0.9600 | 0.0284 | 0.4851 | 0.6096 | 高度耐盐High salt tolerance |
39 | -0.4107 | -0.5841 | 1.7251 | 0.2988 | 0.1245 | 0.8157 | 0.3296 | 中度耐盐Moderate salt tolerance |
40 | -0.0538 | -1.0250 | 0.4973 | 0.3376 | 0.0685 | 0.5648 | 0.2944 | 盐敏感Salt sensitivity |
41 | 6.0342 | 1.7618 | 1.6937 | 1.0000 | 0.4226 | 0.8093 | 0.7993 | 高度耐盐High salt tolerance |
42 | -2.1428 | -0.5681 | 0.0462 | 0.1103 | 0.1265 | 0.4726 | 0.1728 | 高度盐敏感Extremely salt sensitivity |
43 | 0.4245 | 0.1161 | 0.4252 | 0.3897 | 0.2135 | 0.5501 | 0.3632 | 中度耐盐Moderate salt tolerance |
44 | -1.1984 | 0.6045 | 0.4779 | 0.2131 | 0.2755 | 0.5608 | 0.2868 | 盐敏感Salt sensitivity |
45 | -0.9893 | -0.9606 | 0.5596 | 0.2358 | 0.0767 | 0.5775 | 0.2432 | 盐敏感Salt sensitivity |
46 | -2.2146 | -0.6243 | -0.9891 | 0.1025 | 0.1194 | 0.2611 | 0.1327 | 高度盐敏感Extremely salt sensitivity |
47 | 0.5139 | 1.8569 | 0.6272 | 0.3994 | 0.4346 | 0.5913 | 0.4403 | 耐盐Salt-tolerance |
48 | 1.3441 | 0.245 | -0.8058 | 0.4897 | 0.2298 | 0.2986 | 0.3826 | 中度耐盐Moderate salt tolerance |
49 | -0.4474 | 1.3137 | -0.7449 | 0.2948 | 0.3656 | 0.3110 | 0.3183 | 中度耐盐Moderate salt tolerance |
50 | 0.1973 | -0.2087 | 0.3914 | 0.3649 | 0.1722 | 0.5432 | 0.3364 | 中度耐盐Moderate salt tolerance |
51 | 0.6813 | 0.2845 | 0.1753 | 0.4176 | 0.2349 | 0.4990 | 0.3766 | 中度耐盐Moderate salt tolerance |
52 | 0.3868 | 0.2984 | -1.8436 | 0.3856 | 0.2366 | 0.0865 | 0.2940 | 盐敏感Salt sensitivity |
53 | 0.3103 | -0.0096 | -0.0411 | 0.3772 | 0.1975 | 0.4548 | 0.3365 | 中度耐盐Moderate salt tolerance |
54 | -1.3353 | 1.3425 | 0.5085 | 0.1982 | 0.3693 | 0.5671 | 0.3074 | 中度耐盐Moderate salt tolerance |
权重 Index weight(Wj) | 0.5458 | 0.2951 | 0.1591 |
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